Opopaea manongarivo, Daniela & Andriamalala & D.C. & Hormiga & D.C., 2013

Daniela, Andriamalala, D. C., Hormiga, Gustavo & D. C., 2013, Systematics Of The Goblin Spider Genus Opopaea (Araneae, Oonopidae) In Madagascar, Bulletin of the American Museum of Natural History 2013 (380), pp. 1-156 : 110-113

Opopaea manongarivo , new species

Figures 613–647; map 2

TYPES: Male holotype and female allotype from Antsiranana: Réserve spéciale de Manongarivo, 12.8 km 228 ° SW Antanambao, 13 ° 58.69S, 48 ° 25.49E, 780 m, sifted litter, montane rainforest, 11 October 1998, coll. B.L. Fisher (CASC PBI_OON 3847).

ETYMOLOGY: The species epithet is a name in apposition taken from the type locality.

DIAGNOSIS: Males of O. manongarivo can be most easily distinguished from those of other Opopaea species by having a palpal anterior part round, bearing a more or less long, sclerotized embolic extension that folds laterally on the retrolateral side (figs. 613, 629–633, 643, 645, 646). Females of O. manongarivo can be distinguished by having the line uniting the tracheal spiracles sclerotized and the divided parmula with the higher part of the parmula smaller (about 0. 07 mm) and the lower part triangular, larger, and hanging below the lower margin of the scutal ridge measuring about 0.19 mm (figs. 615, 635, 636).

DESCRIPTION: Male (holotype) (PBI_ OON 3847). Small to medium-sized species. Total length 1.45. Color: body orange; legs, palp, and patella yellow. Carapace: pars cephalica elevated, with one row of two widely separated sets of two sharp lateral denticles raised into pointed humps (figs. 626, 627), anteriorly narrowed to two thirds its maximum width. Carapace anterolateral corners without sclerotized, triangular extension. Clypeus low, not rebordered. Chilum present, undivided. Eyes very large. ALE largest. ALE, PME, and PLE oval; PME darkened compared to other eyes, posterior eye row straight viewed from above and from front; ALE separated by less than their radius, ALE-PLE separated by less than ALE radius, PME touching throughout most of their length, PLE-PME separated by less than PME radius. Sternum longer than wide with radial furrows between coxae, surface smooth, without pits, microsculptures only in furrows, shape of posterior part between coxae IV rectangular and swollen (fig. 628). Lateral margins of area between coxae III unmodified, without lumps; setae sparse, dark, needlelike, densest laterally, originating from surface, without hair tufts (figs. 625– 628). Palp cymbiobulbus slightly curved downward, with one posteromedian protuberance. Palpal fenestra large, more or less circular, enlarged anteriorly and located at one third of the palp. Fenestral top flat. Cymbiobulbus almost of the same length and color as patella. Femur longer, inserted at about one third the length from the posterior end of patella (figs. 613, 614, 629, 630, 633, 634, 637–647).

Female (allotype) (PBI_OON 3847). Total length 1.74. Abdomen round and not flattened laterally toward posterior end. Postgynum depression as wide as the distance

between the two lateral apodemes and as long as one third the distance separating the epigynal furrow and the line uniting the tracheal spiracles. Line uniting the tracheal spiracles sclerotized postgynum ridge thinner and barely sclerotized medially (figs. 615, 635, 636).

MEASUREMENTS: Male (holotype) ( PBI _ OON 3847 ). TL: 1.45 ; CL: 0.68; CW: 0.55; CH: 0.31; AL: 0.90; AW: 0.62; ALE: 0.09; PME: 0.09; PLE: 0.08; EGW: 0.24; ALE- ALE: 0.03; ALE-PLE: 0.01; PME-PME: 0; PLE-PME: 0; CBL: 0.26; CBW: 0.09; CBL/ PTL: 1; FI: 0.11.

Female (PBI_OON 3717). TL: 1.74; CL: 0.70; CW: 0.60; AL: 1.06; AW: 0.87; EFL: 0.21; LAL: 0.10; PGI: 0.47; PDL/LAL: 0.50; PDW/EFL: 0.57.

VARIATION: Male: TL: 1.45–1.48; CL: 0.68–0.69; CW: 0.55–0.58; AL: 0.80–0.90; AW: 0.62–0.65.

Female: TL: 1.73–1.74; CL: 0.67–0.70; CW: 0.60–0.61; AL: 1.06–1.06; AW: 0.83– 0.87.

NATURAL HISTORY: O. manongarivo was found in mountain rainforest at high elevation (780–1175 m).

DISTRIBUTION: O. manongarivo was mostly found in the northern and northeastern regions of Madagascar in the province of Antsiranana (Réserve spéciale de Manongarivo) (map 2).

OTHER MATERIAL EXAMINED: MADA- GASCAR: Antsiranana: RS Manongarivo, 14.5 km 220 ° SW Antanambao, S13 ° 59.99, E48 ° 20925.70, 1175 m elevation, 20 October 1998, rainforest, sifted litter, B.L. Fisher (CASC PBI_OON 3717), 28, 1♀ (CASC PBI_OON 2000), 198, 26♀. 12.8 km 228 ° SW Antanambao, 13 ° 58.69S, 48 ° 25.49E, 780 m, sifted litter, montane rainforest, 11 October 1998, coll. B.L. Fisher (CASC. PBI_OON 3847), 48, 8♀.