Olegia brigittae ( Hoernes & Auinger, 1891 ) Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 79-80

Olegia brigittae ( Hoernes & Auinger, 1891) nov. comb.

Figs 25A 1 –A 3 , 1B 1 –B 3 , 5 , 6

Clavatula Brigittae —Hoernes 1891: 129 [nomen nudum].

* Pleurotoma (Clavatula) Brigittae nov. form.— Hoernes & Auinger 1891: 336, pl. 46, figs 8–9.

? [ Pleurotoma ] (Clavatula) Brigittae R. Hörn. & Auinger — Schafarzik 1903: 47

C [lavatula]. (C [lavatula]) brigittae (R. Hörn. et Au.) — Sieber 1958a: 156.

non Clavatula doderleini brigittae Hoernes & Auinger, 1892 — Strausz 1966: 410, pl. 16, fig. 15, pl. 17, fig. 1 [= Olegia mandici nov. sp.].

Type material. Lectotype designated herein: NHMW 2021 /0144/0001, SL: 17.6 mm, MD: 8.0 mm, Lysice (the Czech Republic), illustrated in Hoernes & Auinger (1891, pl. 46, fig. 8), figs 25A 1 –A 3 . Paralectotype: NHMW 2021 /0145/0001, Niederleis ( Austria), SL: 22.7 mm , MD: 9.4 mm, illustrated in Hoernes & Auinger (1891, pl. 46, fig. 9, figs 25B 1 –B 3.

Revised description. Shell small to medium-sized, solid, broad fusiform, with gradate spire; apical angle ~50°. Protoconch not preserved. Teleoconch of at least eight whorls (earliest whorls not preserved). Early teleoconch whorls conical, flat-sided with broad smooth subsutural spiral cord and beaded suprasutural cord, separated by weak concavity. Later whorls subcylindrical; broad, rounded subsutural collar with wide-spaced, poorly defined tubercles, slightly concave, smooth mid-portion, prominent, opisthocline to slightly sigmoidal suprasutural row of tubercles. Suture deeply impressed, weakly undulating. Last whorl 60% of total height. Broad subsutural collar bearing 12–14 indistinct tubercles. Narrow, weakly concave subsutural ramp. Shoulder hardly developed, delimited by row of low, opisthocline, axially elongated tubercles. Almost straight-sided mid-whorl portion bearing subobsolete tubercles. Base weakly constricted. Siphonal fasciole indistinct, rounded, twisted with prominent growth increments. Very weak, irregular spiral cords at mid-whorl and base. Aperture moderately wide, ovate. Outer lip not thickened, smooth within. Anal sinus moderately deep and wide, asymmetrically U-shaped, with apex mid-ramp. Siphonal canal moderately long, narrow. Columella weakly excavated in upper third, straight below, smooth. Columellar and parietal callus adherent, thin, forming indistinct rim.

Discussion. As already recognized by Hoernes & Auinger (1891), Olegia brigittae ( Hoernes & Auinger, 1891) is very close to Olegia baccifera ( Bellardi, 1877) from the Tortonian of Stazzano ( Italy) (see holotype in Bellardi 1877: 166, pl. 5, fig. 29 and Ferrero Mortara et al. 1981: 76, pl. 14, figs 1a–b). A separation of both species is based on the more conical spire of O. baccifera and the more prominent, but less numerous nodes along the abapical suture, and the well-defined nodes on the basal and peribasal cords.

The specimen described by Strausz (1966) from the Pannonian Basin as Clavatula brigittae has spiny nodes, opisthocline axial riblets on early teleoconch whorls and fewer nodes along the abapical suture and is described herein as Olegia mandici nov. sp.

Paleoenvironment. At the Niederleis section, sediment and fossils from coastal and lagoonal environments not exceeding 30 m water depth were transported by tempestites into offshore settings (Mandic et al. 2002). No information is available for the Lysice section.

Distribution in Central Paratethys. Badenian (middle Miocene): Alpine-Carpathian Foreland Basin: Lysice (the Czech Republic); Vienna Basin: Niederleis ( Austria) ( Hoernes & Auinger 1891). A further occurrence was mentioned by Schafarzik (1903) from Budapest ( Hungary), but we have not seen this material to verify the identification.

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 6. Apertures of Paratethyan Clavatulidae (C. = Clavatula, ‘C.’ = ‘Clavatula’, G. = Granulatocincta, M. = Megaclavaltula, O. = Olegia).

FIGURE 25. A1–A3. Olegia brigittae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0144/0001, Lysice (the Czech Republic). B1–B3. Olegia brigittae (Hoernes & Auinger, 1891), paralectotype, NHMW 2021/0145/0001, Niederleis (Austria). C1–C3. Olegia dorotheae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0040a, Kleinebersdorf (Austria). D1–D3. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). E1–E3. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW NHMW 1949/0005/0040c, Kleinebersdorf (Austria). F1–F3. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 2021/0146/0001, Kleinebersdorf (Austria).