Onthophagus (Onthophagus) dicranoides Balthasar, 1939

Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z., 2019, Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records, ZooKeys 826, pp. 1-343 : 183

Onthophagus (Onthophagus) dicranoides Balthasar, 1939 Plate 41A

Onthophagus dicranoides Balthasar, 1939h: 43 (original description. Type locality: Bucai [= Bucay], Guayaquil).

Onthophagus dicranoides : Martínez 1947: 112 (distribution); Zunino and Halffter 1997: 165 (list of species); Pulido-Herrera and Zunino 2007: 102 (catalog of species); Carvajal et al. 2011: 322-323 (cited for Ecuador); Krajcik 2012: 178 (complete list of species); Bezdek and Hajek 2013: 404 (catalog of the types of the NMPC).

Onthophagus (Onthophagus) dicranoides : Chamorro et al. 2018: 97 (cited for Ecuador).

Type specimens.

Onthophagus dicranoides Balthasar, 1939. Five syntypes examined deposited at the NMPC and MSMF. Lectotype to be designated in a future work on this species group.

Syntype (♂): "W Ecuador / Pucay / F. Ohaus S. [p]", "Bucay 300 m / F. Ohs. 12.6.05 [p]", "P.G. 3345 / Canada balsam / M. Zunino 1980 [hw]", "TYPUS [p, red label, black margin]", "Onthophagus / dicranioides / Dr. V. Balthasar det. [p and hw]", "Mus. Nat. Pragae / 26326 / Inv. [p and hw, orange label]", "dicranioides m. [hw, green label, black margin]".

Syntype (♀): "Bucay 300 m / F. Ohs. 12.6.05 [p]", "P.G. 3346 / Canada balsam / M. Zunino 1980 [hw]", "TYPUS [p, red label, black margin]", "Onthophagus / dicranioides / n. sp / Dr. V. Balthasar det. [p and hw]", "Mus. Nat. Pragae / 26327 / Inv. [p and hw, orange label]".

Syntype (♀): "Bucay 300 m / F. Ohs. 20.6.05 [p]", "Onthophagus / dicranioides / n. sp Typ. / Dr. V. Balthasar det. [p and hw]", "Senckenberg- / Museum / Frankfurt / Main [p]", "TYPUS [p, red label, black margin]", "Senckenberg / Museum [p]".

Syntype (♂): "Bucay 300 m / F. Ohs. 20.6.05 [p]", "Onthophagus / dicranioides / n. sp Typ. / Dr. V. Balthasar det. [p and hw]", "Senckenberg- / Museum / Frankfurt / Main [p]", "TYPUS [p, red label, black margin]", "Senckenberg / Museum [p]".

Syntype (♂): "W Ecuador / Guayaquil [p]", "Onthophagus / dicranioides / n. sp. / Dr. V. Balthasar det. [p and hw]", "Senckenberg- / Museum / Frankfurt / Main [p]", "Para- / typoid [p, black label]".

Distribution.

Only known from Ecuador.

Records examined.

CAÑAR: La Troncal (2 specimens CEMT). EL ORO: Piñas, 1200 m (9 specimens CEMT). GUAYAS: Pucay [= Bucay], 300 m (2 specimens NMPC; 2 specimens MSMF); Guayaquil (1 specimen MSMF). LOS RÍOS: CCRP [= Centro Científico Río Palenque] (13 specimens CEMT); Río Palenque Station (1 specimen CEMT). SANTO DOMINGO DE LOS TSÁCHILAS: Santo Domingo de los Colorados (1 specimen CEMT).

Temporal data.

Collected in January, February, March, May, June, and September.

Remarks.

Inhabits coastal lowland evergreen forests and evergreen foothill forests from 50-1200 m a.s.l. The collection method is unknown.