Paramannokeraia juliae van Achterberg

Achterberg, Cornelis van, Quicke, Donald L. J. & Boring, C. Andrew, 2017, A revision of the tribe Planitorini van Achterberg (Hymenoptera, Braconidae, Euphorinae), with description of a new genus from Australia, ZooKeys 718, pp. 35-64 : 46-48

Paramannokeraia juliae van Achterberg sp. n. Figs 66, 67-76, 77-83

Mannokeraia sp.; Stigenberg et al. 2015: 575.

Type material.

Holotype, ♀ (NHRS), "Australia, Tasmania, Cradle Mtn NP, creek from Crater Lake to Ronny Creek, 100 m upstr. boardwalk, 867 mao, S41°38.667' E145°56.755', 23. ii– 4.iii.2006, Malaise trap, loc. 14, N. Jönsson, T. Malm & D. Williams", "DNA voucher DNA JS10_00282", " NHRS-HEVA 000004017". Paratype: 1 ♂ (CNC), "Australia, Tas[mania], Mt. Field NP, 7.i.1984, L. Masner, s. s.".

Diagnosis.

Antenna of ♀ with 19 robust segments, apical segments pedunculate (Fig. 72), of ♂ cylindrical and elongate (Fig. 81); head transverse, not enlarged behind eyes in dorsal view (Fig. 74); face mainly sparsely punctate (Fig. 73); mesosoma of ♀ normal, with mesoscutum distinctly above upper level of pronotum (Fig. 68); notauli on mesoscutal disk widened (Fig. 69); propodeum rather sparsely rugulose but sublaterally largely smooth (Fig. 69); base of hind coxa smooth dorsally (Fig. 68); hind femur rather swollen (Fig. 71); fore tibia rounded apically, without tooth-like protuberance apically (Fig. 76); fore and middle tarsi of ♀ widened (Figs 66, 76); basal half of antenna of both sexes and legs dark brown; setose part of ovipositor sheath about 0.4 times as long as hind tibia (Fig. 66); both sexes macropterous.

Description.

Holotype, ♀, length of fore wing 2.5 mm, and of body 2.7 mm.

Head. Antenna with 19 segments, length of third segment 1.1 times fourth segment, third, fourth and penultimate segments 1.6, 1.4 and 0.9 (without pedunculus 0.8) times as long as wide, respectively (Fig. 72) and with apical 12 segments pedunculate, medially antenna as wide as apically; length of maxillary palp 0.8 times height of head; occipital carina complete, low dorsally (Fig. 75); eye 1.5 times as long as temple in dorsal view; temples subparallel-sided behind eyes; OOL:diameter of posterior ocellus:POL = 9:5:11; frons smooth and shiny, with some long setae, convex, with shallow median groove, and anteriorly flattened; face with some rugae dorsally below antennal sockets and remainder sparsely coarsely punctate (Fig. 73); clypeus depressed and smooth ventrally, with ventral rim slightly upcurved, dorsally weakly convex and with some coarse punctures (Fig. 73); length of malar space 1.1 times basal width of mandible; occipital carina joining hypostomal carina and occipital flange subcircular; mandible depressed medially and shiny, apically with large upper and small lower tooth.

Mesosoma. Length of mesosoma 1.9 times its height; dorsal pronope and antescutal depression absent; side of pronotum largely punctate-rugose ventrally, largely smooth medially, with narrow crenulate groove antero-dorsally and punctate-costate posteriorly; epicnemial area punctate dorsally; precoxal sulcus complete, narrowly crenulate-punctate (Fig. 68); remainder of mesopleuron smooth; mesosternal suture deep and coarsely crenulate; postpectal carina absent; notauli complete, coarsely punctate and rather wide, ending in wide punctate area (Fig. 69); scutellar sulcus with five costae; scutellum flat, smooth and shiny; metapleuron punctate medially and coarsely reticulate-punctate ventrally; postpectal carina absent and area between middle coxae with few punctures; propodeum rather sparsely rugulose but sublaterally largely smooth (Fig. 69), posterior face weakly differentiated and without an obtuse tubercle postero-laterally (Fig. 68).

Wings. Fore wing: 1-M weakly curved; 1-SR short (Fig. 67); marginal cell open anteriorly because of most of 1-R1 absent and sclerotized part of 1-R1 about 0.2 times as long as pterostigma (Fig. 67); vein r emitted distinctly after middle of pterostigma; r:3-SR:SR1 = 2:11:62; vein SR1 straight; 2-SR:3-SR:r-m = 27:11:15; 2-M much longer than 3-SR; m-cu slightly postfurcal; 1-CU1 oblique and narrow, about as long as cu-a; 1-CU1:2-CU1 = 5:27; basal and subbasal cells of fore wing setose as other cells. Hind wing: marginal cell subparallel-sided medially and obsolescent apically; M+CU:1-M:1r-m = 41:18:10.

Legs. Hind coxa basally smooth; tarsal claws with wide truncate lamelliform lobe (Fig. 71); length of femur, tibia and basitarsus of hind leg 3.0, 7.3 and 6.0 times as long as their maximum width; fore femur rather inflated, 2.9 times longer than wide and apically rounded (Fig. 76); fore and middle tarsi widened (Figs 66, 76).

Metasoma. First tergite 1.5 times longer than its apical width, distinctly petiolate (Fig. 70), with incomplete straight striae, dorsal carinae unite to form a median carina and dorsope deep and large (Fig. 70), only basal quarter closed ventrally; laterope absent, tergite widened latero-basally; second tergite smooth; ovipositor sheath somewhat widened and obtuse apically (Fig. 66), its setose part 0.14 times as long as fore wing and 0.40 times hind tibia; ovipositor with minute subapical nodus and widened basally.

Colour. Black; antenna, metasoma except black first tergite and legs dark brown, but hind trochanter and tibial spurs brown; tegulae and palpi pale yellowish; pterostigma and veins brown; wing membrane weakly infuscate.

Male. Similar to female holotype except for the shape of the antennal segments, slender fore and middle tarsi (Figs 77, 80, the rugose area between middle coxae and the different sculpture of the propodeum and face (Figs 78, 82). Antenna with 28 segments, length of fore wing 3.6 mm, and of body 3.6 mm; face and clypeus rather finely punctate; metasoma (except most of first tergite) brown; mesoscutum less flattened; medio-posterior punctate area of mesoscutum small; propodeum largely finely rugulose; first tergite 1.5 times longer than wide apically and distinctly longitudinally striate.

Etymology.

Named after Dr Julia Stigenberg (Stockholm), who generously made the holotype available for this study.

Distribution.

Australia (Tasmania). Collected in January–March.