Phagnalon methanaeum Haussknecht (1887: 85)

Montes-Moreno, Noemí, Garcia-Jacas, Núria, Nualart, Neus, Susanna, Alfonso & Sáez, Llorenç, 2018, Typification of plant names referable to Phagnalon (Compositae) with some taxonomic notes, Phytotaxa 360 (1), pp. 1-18 : 8

Phagnalon methanaeum Haussknecht (1887: 85)

Type:—[ GREECE.] Methana , 25 May 1885, Haussknecht s.n. (lectotype: JE barcode JE00017037 [digital image!], designated here; isolectotypes: BM barcode BM001025787 !, K barcode K000910410 !) .

(Image available at: https://plants.jstor.org/stable/10.5555/al.ap.specimen.je00017037).

= Phagnalon × intermedium (Lag.) Pau (1931: 51)

Notes:—The locality indicated in the original description of Ph. methanaeum is: “Das in Griechenland sehr häufige Ph. graecum B. weicht durch schmale und spitze Hüllschuppen sehr ab. an Kalkfelsen der Halbinsel Methana bei Vromolimni”. We found two specimens corresponding to the original material at JE, where Haussknecht deposited his main collection, as well as other herbaria (BM and K). We selected the specimen that better agrees with the protologue. It bears a note “n. sp.” and a handwritten label with the collection date. We also found specimens of the original gathering collected in the exact original locality at K and BM, but we did not choose any of them as the lectotype because the note “n. sp.” was not written on the label. Similarly, other specimens collected from the original locality and labelled as type material are deposited at BR, C, K, LD and M. However, these specimens were collected after the protologue was published and they should not be considered as part of the original material. The protologue indicated that Ph. methanaeum is close to Ph. rupestre , but our study of the original material deposited at JE revealed that these specimens show intermediate characters attributed to hybridization between Ph. rupestre and Ph. saxatile . They also have linear to lanceolate leaves, like those of Ph. saxatile , the outermost and middle involucral bracts being wider and obtuse, like those of Ph. rupestre , but frequently with undulate margins, like those of Ph. saxatile .