Prosyntexis gouleti Sharkey, 1990

Jattiot, Romain, Krogmann, Lars & Nel, André, 2011, Revision of Prosyntexis from the Lower Cretaceous Crato Formation of Brazil (Hymenoptera: Sepulcidae: Trematothoracinae), Zootaxa 3058 (1), pp. 55-62 : 56-61

Prosyntexis gouleti Sharkey, 1990

( Figures 1–10 )

Material. Specimen MNHN.F. A43720 (coll. Borschukewitz), deposited in the Laboratoire de Paléontologie, Muséum National d’Histoire Naturelle, Paris, France; Specimen SMNS 66301 and specimen SMNS 66698a (imprint) – 66698b (counterimprint), deposited in the fossil collection of the State Museum of Natural History Stuttgart, Germany.

Age and outcrop. Lower Cretacous Crato Formation of Northeastern Brazil.

Description. Specimen MNHN.F. A43720 ( Figs 1–2 ): Body visible from below, 13.0 mm long (14.5 mm with ovipositor); head 1.2 mm long, 2.0 mm wide; antenna not preserved; thorax 5.5 mm long, 3.0 mm wide; abdomen 5.5 mm long, 2.0 mm wide; ovipositor sheath large, 5 mm long, 1.4 mm wide; ovipositor 1.7 mm long, 0.8 mm wide; six legs visible but poorly preserved. Wings without trace of coloured spots (maybe originally hyaline or uniformly coloured); fore wing 10.5 mm long, 4.0 mm wide, pterostigma brown, rather broad, 2.5 mm long, 0.5 mm wide, with its base 5.5 mm from wing base; subcosta completely fused with R; area between costal margin and R very narrow, 0.2 mm wide; Rs emerging from R 3.3 mm from wing base, with basal portion slightly curved, 1.1 mm long, rather long before its fusion with M; M+Cu basally curved and straight in its distal part; M and Cu separating 3.8 mm from wing base; basal free part of M short, 0.5 mm long; straight Rs+M 0.9 mm long; distal Rs+M forking into Rs and M slightly before 1m-cu; distal part of Rs between Rs+M and 1r-rs straight, 0.6 mm long; 1r-rs 0.6 mm long; 2r-rs 0.4 mm long, emerging from pterostigma in its basal half, 1.0 mm from its base; 2r-m absent but 3r-m clearly present and slightly curved, 0.8 mm long; cell 1R1 elongate, 2.3 mm long, 1.1 mm wide; cell 2R1 0.9 mm long, 0.65 mm wide; cell 3R1 3.0 mm long, 1.0 mm wide; cell Rs(+1Rs2) 2.8 mm long, 0.85 mm wide; cell 1M 1.1 mm long, 0.6 mm wide; cell 2M 2.4 mm long, 1.2 mm wide; crossvein cu-a present just distal of base of free part of M, 0.5 mm long; cell 1Cu 3.5 mm long; cell 2Cu larger and broader than cell 1M, 1.3 mm long, 0.8 mm wide; cell 3Cu 2.8 mm long; crossvein a present at 2.5 mm from wing base; basal part of 2A present; vein 1A fused at apex with vein 2A+3A; cell 1A 2.6 mm long, cell 2A 1.8mm long. Hindwing only partly preserved, probably hyaline or uniformly coloured, with apico-costal part preserved, basal part of Rs very long, 2.0 mm long, distal part of Rs 1.9 mm long, cell R1 closed, 3.8 mm long, 0.6 mm wide; 1r-m 0.6 mm long, basal part of M 1.1 mm long, no crossveins 2r-m or 3r-m.

Specimen SMNS 66301 ( Figs 4–5 ): Total body length 10.3 mm (9.44 mm without ovipositor). Head 1.17 mm long, 1.66 mm wide; antennae nearly completely preserved (apart from proximal portion); limits of scape and pedicel indiscernible; first flagellomere 0.69 mm long, 0.20 mm wide; second flagellomere 1.21 mm long, 0.18 mm wide; third flagellomere 1.10 mm long, 0.18 mm wide; limits of remaining flagellomeres not clearly indicated, total number of antennomeres 13 – 15 ( Fig. 5 ). Thorax 2.60 mm long, 1.34 mm wide; metascutellum 0.29 mm long, 0.46 mm wide. Wing venation ( Fig. 4 ) identical to that of specimen MNHN.F. A43720 . Abdomen 5.89 mm long, 2.50 mm wide; ovipositor 0.86 mm long, 0.47 mm wide.

Specimen SMNS66698 ( Figs 6–10 ): Imprint (SMNS 66698a, Figs 6 – 7 ) and counterimprint (SMNS 66698b, Figs 8 – 10 ) preserved. Total body length 11.0 mm. Head 1.52 mm long, 1.66 mm wide, eyes large, ocelli partly preserved in counterimprint ( Fig. 9 ); antennae incompletely preserved; first flagellomere 0.74 mm long, 0.21 mm wide; remaining flagellomeres indiscernible. Posterior limit of thorax indiscernible, thorax 1.96 mm wide. Wing venation identical to that of specimen MNHN.F. A43720 ( Figs 7 , 10 ). Anterior limit of abdomen indiscernible, abdomen 2.60 mm wide; ovipositor not distinct.

Figures

FIGURES 1–3. Prosyntexis gouleti Sharkey, 1990. (1) Specimen A43720, photograph of right forewing (scale bar: 2 mm); (2) specimen A43720, photograph of left forewing (scale bar: 2 mm); (3) holotype specimen of ‘Prosyntexis legitima’ (syn. nov.), drawing of forewing venation after the original photograph of Martins-Neto et al. (2007: fig. 2A), wing cells in bold, veins in normal font.

FIGURES 4–5. Prosyntexis gouleti Sharkey, 1990, specimen SMNS 66301. (4) Photograph of habitus in dorsal view (scale bar: 5 mm); (5) photograph of antennae (scale bar: 1 mm), with numbers of antennomeres indicated.

FIGURES 6–7. Prosyntexis gouleti Sharkey, 1990, specimen SMNS 66698a, imprint. (6) Photograph of habitus in dorsal view (scale bar: 5 mm); (7) photograph of left forewing (scale bar: 2 mm).

FIGURES 8–10. Prosyntexis gouleti Sharkey, 1990, specimen SMNS 66698b, counterimprint. (8) Photograph of habitus in ventral view (scale bar: 5 mm); (9) photograph of head (scale bar: 200 µm); (10) photograph of right wings (scale bar: 2 mm).