Rhyacophila arefini Lukyanchenko 1993

Ito, Tomiko, Kuhara, Naotoshi & Malicky, Hans, 2024, Five new species of the genus Rhyacophila Stephens (Trichoptera, Rhyacophilidae) from southwestern Hokkaido, northern Japan, Zootaxa 5536 (4), pp. 551-568 : 552

Rhyacophila arefini Lukyanchenko 1993

( Figs 1A–1C )

Rhyacophila arefini Lukyanchenko 1993, pp. 5–6 , male, female, pupa, larva, South Kuril (Kunashir), Hokkaido (Ishikari); Arefina 2001, pp. 28, 41, female, South Kuril (Shikotan); Hattori 2005, pp. 427, 434–435, male, female, Hokkaido.

Records in references. Kuhara & Ito 1994, pp. 20, 23, Hokkaido ( Ishikari ) ; Vshivkova & Kolin 1997, p. 60, Sakhalin ; Ito et al. 1998, pp. 10, 12–14, 16, 17, Hokkaido ( Nemuro ) ; Ohkawa 1999, pp. 40–44, Hokkaido ( Tokachi ) ; Kuhara & Sato 2001, p. 64, Hokkaido ( Rishiri Island ) ; Ohkawa & Hattori 2007, p. 266, Hokkaido ( Kamikawa ) ; Muramatsu 2008, p. 3, Hokkaido ( Okhotsk ) ; Ito et al. 2009, p. 8, Hokkaido ( Okhotsk ) ; Ito & Kubo 2011, p. 4, Hokkaido ( Tokachi ) ; Kuhara 2011, pp. 52, 57, 61, Hokkaido (Sorachi, Hidaka ) ; Iijima & Tsuchiya 2014, p. 27, Hokkaido ( Okhotsk ) ; Ito & Kuhara, 2023, p. 63, Hokkaido (Soya, Kamikawa ) ; Ito et al. in press, Sakhalin .

Material examined. Hokkaido, Sorachi: Akabira-shi , Penke-shubushinai-zawa, 67 males, 38 females, 29.vi– 12.vii.2016, leg. Y. Nagasaka et al., M ( TI). Ashibetsu-shi, Nakanosawa-gawa, 9 males, 13.vi–8.viii.2013, leg. Y. Nagasaka et al., M ( TI). Ashibetsu-shi, Nisen-zawa, 1 male, 28.vi–12.vii.2013, leg. Y. Nagasaka et al., M ( TI). Ashibetsu-shi, Yoshida-no-sawa, 2 males, 1 female, 7.vi–23.viii.2013, leg. Y. Nagasaka et al., M ( TI). Ishikari: Sapporo-shi, Takino, Masuno-sawa, small tributary, 1 male, 23.vii.1977, leg. TI , S ( TI); 3 males, 6.vi–18.vii.1981, leg. TI , S ( TI); 1 male, 22.vi.1984, leg. TI , S ( TI); 1 male, 18.vii.1986, leg. TI , S ( TI). Sapporo-shi, Jôzankei, Yoichidake , 1,200 m, 1 male, 8.viii.1982, leg. TI , S ( TI). Eniwa-shi, Moizari-gawa, Saiwai-bashi , 5 males, 7 females, 5– 10.vi.2013, leg. TI , L ( TI). Eniwa-shi, Kita-kashiwagi, Fishery Experimental Station , 2 males, 2 females, 28.v.1996, leg. TI , S ( TI). Eniwa-shi, Eniwa-kôen , 2 males, 1 female, 19.vii.1996, leg. TI , S ( TI). Chitose-shi, Rankoshi , salmon hatchery, sprimg stream, 1 male, 5.vi.1980, leg. TI , S ( TI). Chitose-shi, Rankoshi, spring stream near Fukajô-bashi , 5 males, 4 females, 5.vi.1980, leg. TI , S ( TI); 2 males, 9.vii.2004, leg. TI , S ( TI); 1 male, 6.vii.2017, leg. TI , S ( TI).

Distribution ( Figs 1A–1C , black circles). Hokkaido (northern, eastern and central areas), South Kuril, Sakhalin.

Habitat. The adults of the species were collected in mountain areas near springs, waterfalls, and small streams up to several meters wide.

Species group. This species was placed in the Rhyacophila sibirica Species Group ( Lukyanchenko 1993).

Japanese name. Arefin-nagare-tobikera.

Figures

FIGURE 1. Recorded collection sites of Rhyacophila species. 1A, location of Hokkaido and recorded sites of known species discussed in present study. 1B–1E, recorded and collected sites in Hokkaido: 1B, 1C, R. eniwa sp. nov. and R. arefini Lukyanchenko; 1D, R. bifue sp. nov., R. imakane sp. nov., R. misago sp. nov., and R. hamata Hattori & Ohkawa; 1E, R. shimamaki sp. nov.