Schismatogobius sapoliensis, Keith & Dahruddin & Limmon & Hubert, 2018
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publication ID |
https://doi.org/ 10.26028/cybium/2018-422-007 |
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persistent identifier |
https://treatment.plazi.org/id/9C0CC15D-CA06-196D-C1EB-FDF823B5F8CB |
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treatment provided by |
Felipe |
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scientific name |
Schismatogobius sapoliensis |
| status |
sp. nov. |
Schismatogobius sapoliensis , n. sp.
( Figs 1-3 View Figure 1 View Figure 2 View Figure 3 ; Tabs I-II)
Material examined
Five specimens from Halmahera with a size range of 16.2-18 mm SL.
Holotype. – MZB 24586 View Materials , male (17.9 mm SL); Samuda , Sapoli , Halmahera, Indonesia, 23 March 2017, Hubert et al. coll.; BIF 6802.
Paratypes. – MZB 24587 View Materials , 2 females (16.2-17.6 mm SL); same data as for holotype; BIF 6803 & 6805 . MNHN 2016- 0626 About MNHN , 1 female (18 mm SL); same data as for holotype; BIF 6801 . MNHN 2016-0627 About MNHN , 1 male (17.8 mm SL); same data as for holotype; BIF 6804 .
Diagnosis
Usually 16 pectoral rays; pectoral fins with a large transverse medium black band. First dorsal fin membrane posterior to spine 6 not connected to base of spine of second dorsal fin. Ventral surface of head in male blackish or greyish with a white mentum. Frenum whitish with a blackish border. Ventral surface of head in female blackish or greyish, except the isthmus, which is whitish, and with a white mentum. Frenum whitish, sometimes with a distal black dot. A single mitochondrial lineage was observed for this species (BOLD: ADL4589).
Description
A small sized Schismatogobius (average adult size <20 mm SL). Body naked, slender, almost circular in cross-section. Head rounded, snout rather pointed. Mouth oblique, lower lip more prominent. Jaw length in male much greater than in female; jaw length 47.4-52.8% of HL in males and 32-36.7% of HL in females. Lower jaw reaching vertical of 1/3 to 1/2 of the eye in female and exceeding a vertical of posterior margin of eye in male. Eyes high on head, close together with interorbital width about equal to 1/3 eye diameter. Anterior nostril short and tube-like.
Dorsal fins VI-I,8-9, membrane in first dorsal fin posteri- or to spine 6 not connected to base of spine of second dorsal fin. D1 with all spines about equal in length. Anal fin I,8-9, origin directly opposite to second dorsal fin origin. Caudal fin with 10-11 branched rays, posterior margin straight. Pectoral fins oblong with posterior margin rounded and 16(4)-17(1) rays, ventralmost ray unbranched. Pelvic fins always I,5, with both fins joined together for their entire length between fifth rays to form a strong cup-like disc and a well-developed and lobed frenum between spines; fins not extending beyond anus. Morphomeristic data given in table II.
Tongue bilobed. Teeth in upper jaw (first row 8-10) in two rows, teeth conical and slightly recurved. Teeth in lower jaw (first row 12-13) in 2-3 rows, all teeth conical with outer row teeth only slightly enlarged and somewhat recurved.
Cephalic sensory pore system always with pores B, D, F, K, L, N and O, pore D singular with all other pores paired; oculoscapular canal absent between pores F and K. Anterior interorbital extension of anterior oculoscapular canal with double terminal pores B slightly posterior to posterior nostril. D pore at rear of interorbital. Posterior extension of anterior oculoscapular canal terminating laterally on each side of head at pore F, just behind posterior edge of eye. Posterior oculoscapular canal with 2 terminal pores, K and L; preopercular canal with 2 pores, N and O. Cutaneous sensory papillae not well developed.
Sexual dimorphism fairly well developed with male having jaws longer than female and a different colour pattern on ventral surface of head. Urogenital papilla oval in female and slightly pointed in male.
Colour in preservation
Body with three or four vertical black bands in dorsal view. When four bands, first band below first dorsal fin, second and third bands below second dorsal fin (sometimes these two bands are overlap) and fourth band at hypural crease. These lateral black body markings alternate with 3 vertical white to grey stripes. Head dusky grey. Ventral surface of head in male blackish or greyish with a white mentum. Frenum whitish with a blackish distal border. Ventral surface of head in female blackish or greyish except the isthmus, which is whitish, and with a white mentum. Frenum whitish, sometimes with a distal black dot ( Fig. 2 View Figure 2 ). First dorsal fin with a large horizontal black band. Second dorsal fin mostly cream with rows of black spots on rays. Caudal fin black and white, with a black spot at centre of hypural crease and two white spots posteriorly. Anal fin mostly greyish. Pectoral fins greyish with a large transverse medium black band.
Colour in life ( Figs 3 View Figure 3 , 4 View Figure 4 )
Three or four vertical black bands in dorsal view. When four bands, first band below first dorsal fin, second and third bands below second dorsal fin and fourth band at hypural crease; the two middle bands are very close to each other. These bands are not uniform but with dorsal or lateral brown to rose spots. Colour of body between each band rose to greyish. Head and cheeks usually mottled with closely spaced black spots and markings, and an overall shade of greyishbrown. Ventral surface of head in male blackish or greyish with a white mentum. Frenum whitish with a blackish border. Ventral surface of head in female blackish or greyish except the isthmus, which is whitish, and with a white mentum. Frenum whitish, sometimes with a distal black dot. Belly whitish to greyish. First dorsal fin with a large horizontal black band. Second dorsal fin with rows of black spots on rays. Caudal fin mainly black with two median white spots dorsally and ventrally. Pectoral fins translucent with a large transverse medium black band.
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