Tipula (Yamatotipula) stackelbergi Alexander

Salmela, Jukka, 2012, Revision of Tipula (Yamatotipula) stackelbergi Alexander (Diptera, Tipulidae), and a short discussion on subspecies among crane flies, ZooKeys 162, pp. 43-58 : 45-46

Tipula (Yamatotipula) stackelbergi Alexander stat. rev. Figs 123e6a, c, d

Tipula (Tipula) stackelbergi Alexander 1934: 305.

Tipula (Yamatotipula) pruinosa stackelbergi Savchenko 1961: 292.

Tipula (Yamatotipula) pruinosa stackelbergi Oosterbroek and Theowald 1992: 165.

Tipula (Yamatotipula) pruinosa stackelbergi Oosterbroek 2011: http://nlbif.eti.uva.nl/ccw/

Material examined.

Holotype of Tipula stackelbergi : male, pinned specimen (ZISP). "Tigrovaja, Suchan./ rn.Uss.kr. 16.VI/ Stackelberg. 927" (white label, partly hand written, in Cyrillic letters). “81” (white label, handwritten). "HOLOTYPE/ Tipula stackelbergi/ C.P. Alexander" (red label, partly handwritten) (Fig. 1a).

With except of the male hypopygium, the holotype specimen is in rather good condition (Figs 1b, c). All legs are detached from the specimen, but four legs are glued to the pin below the specimen. Tips of wings are broken. Right antenna is broken, only scape and pedicel are left; left antenna has seven flagellomeres. Tip of abdomen is broken; apparently hypopygium is mounted on a celluloid strip, which is attached on a pin. The surface of this strip is heavily cracked, and the structure of the hypopygium cannot be examined.

Other material.

Russia, Vladivostok, Nekrutenko leg, 2.VI. 1957, 1 male, 1 female (ZISP).

Redescription.

Male. Head yellowish brown, with grey pruinosity. Rostrum yellowish, nasus distinct, bearing numerous light hairs. Palpi brown. Scape yellowish, elongate, length 387-450 μm, width 126-131 μm (n=2). Pedicel yellowish, globular, length 147 μm, width 139 μm (n=1). Flagellomere 1 yellowish brown, length 486 μm, width 91 μm (n=1). Flagellomere 2 length 464 μm, width 79 μm (n=1). Flagellomeres bear erect short hairs, giving silvery appearance. Flagellomeres 2-7 elongate, brown, with dark verticils (Fig. 1c).

Prescutum with four brown stripes (Fig. 1e). Pronotum, prescutum, scutum, anepisternum, katepisternum and meron brownish, with grey pruinosity. Scutellum, anepimeron and laterotergite yellowish. Anterior part of mediotergite yellowish, more brownish in posterior part, having two weak longitudinal brown stripes. Coxa 1 brown. Anterior part of coxa 2 brown, posterior part yellow. Coxa 3 yellow. Femorae yellowish brown, darkening toward tarsi. Wings without markings, pterostigma brown (Fig. 1f). Wing length 13.8 mm (n=1). Halter yellowish.

Abdominal tergites yellowish brown, slightly darkening toward tip of abdomen. 9th tergite with two median projections, densely covered by dark bristles. Lateral corners of 9th tergite glabrous, pointed (Fig. 2a). 9th sternite with median incision, bearing two fleshy and hairy outgrowths in the margin of the incision. Outer gonostylus worm-like, apical half covered by light hairs (Fig. 2b). Inner gonostylus elongate (Figs 2b, c, 3e); beak rounded, with ten stout apical bristles and four subapical weaker bristles; central ridge with few weak bristles along its length; lower beak roundish, not angular. Posterior immovable apodeme of sperm pump almost straight (Fig. 2d). Distal end of compressor apodeme of sperm pump club-shaped, roundish (Fig. 2f). Aedeagal guide as in Fig. 2e.

Female. In general similar to male. Scutellum brown, abdominal tergites brown. Wing length 18.4 mm (n=1). Female terminalia as in Fig. 6a. Basal part of hypogynial valves with dense black setae. Proximal ends of valves roundish, tapering toward bases (Fig. 6c). Genital fork of vaginal apodeme brown, rather narrow in its whole length (Fig. 6d). Dorsal view of vaginal apodeme as in Fig. 6d.