Tragardhus (Tragardhus) majae, Kamiński, 2017

Kamiński, Marcin Jan, 2017, Afrotropical Melambiina: revision of the genus Tragardhus Koch, 1956 and redescription of Pseudemmallus Koch, 1956 (Tenebrionidae: Pedinini), Zootaxa 4347 (1), pp. 71-93 : 85-86

Tragardhus (Tragardhus) majae sp. nov.

http://zoobANk.oRg/URN:lsiD:zoobANk.oRg:Act:274B8B4D-09F1-45BA-A653-15C5E9726A76 ( Figs 4D , 7 )

Studied type material. Holotype, mAle ( SMNS): “ RSA: KwAZUlU NAtAl / 15 km E VRYheit / 26.I.2008 / leg. P. Schüle ” . Paratype, femAle ( SMNS): “ RSA, KwAZUlU-NAtAl / VRYheiD 29/ 31.12.2008 / P. Schüle leg.” ( Fig. 4D ). Etymology. NAmeD AfteR MAjA LAURA Szczepłek (boRN oN FebRUARY 2 ND, 2016), DAUghteR of mY fRieNDs DomiNikA GóRAlczYk AND JAN Szczepłek.

Diagnosis. See DiAgNosis of T. (T.) jani sp. nov.

Description. BoDY—leNgth = 4.5–5.0 mm. HAbitUs As iN Fig. 4D .

DoRsAl siDe of heAD DUll, with coNspicUoUs pUNctURes (0.1 DiAmeteRs ApARt). FRoNtoclYpeAl sUtURe NoticeAble; with pAiR of Deep DepRessioNs locAteD NeAR meDiAN pARt. CANthUs sUbpARAllel At bAse. ClYpeAl emARgiNAtioN Deep (clYpeAl emARgiNAtioN wiDth: Depth RAtio cA. 5.0). LAbRUm wiDe (wiDth: leNgth RAtio cA. 2.9); DelicAtelY emARgiNAte iN miDDle; posteRioR thiRD coveReD with DeNse AND coNspicUoUs setAe. MeNtUm wiDest iN miDDle; flAt, Not coveRiNg cARDo oR stipes; ApicAllY with shAllow meDiAN iNDeNtAtioN; lAteRAl lobes, iN veNtRAl view, coveReD bY miDDle pARt. SUbmeNtUm tRiANgUlAR. ApicAl segmeNt of mAxillARY pAlpUs ovAl; with seNsoRY fielD occUpYiNg whole pRoximAl eDge. SecoND segmeNt of mAxillARY pAlpUs eloNgAteD (leNgth: wiDth RAtio cA. 3.8). PAlpifeR with pAtch of loNg setAe locAteD NeAR bAse; with iNteRNAl sUlcUs foR mAxillARY pAlpUs visible fRom veNtRAl view. ANteRioR teNtoRiAl pit ciRcUlAR. EYe DoRsAllY shielDeD bY loNgitUDiNAl keel. DoRsAl AND veNtRAl pARts of eYes composeD of similAR NUmbeR of fAcets; thRee fAcets pReseNt betweeN cANthUs AND temple. ANteNNA RelAtivelY loNg (ANteNNA: pRoNotUm leNgth RAtio cA. 1.2); thiRD ANteNNomeRe shoRt (3RD ANteNNomeRe: 2ND ANteNNomeRe leNgth RAtio cA. 2.0); fifth ANteNNomeRe shoRt (leNgth: wiDth RAtio cA. 1.1); ANteNNomeRes 8–10 tRANsveRse.

PRoNotUm NARRoweR thAN elYtRA (elYtRA: pRoNotUm wiDth RAtio cA. 1.2); RelAtivelY wiDe (leNgth: wiDth RAtio cA. 0.9); Disc DUll, coveReD with coARse pUNctURes (0.1 DiAmeteRs ApARt); with two pAiRs of RoUNDeD cARiNAe; ANteRioR ANgles shARp AND stRoNglY pRotRUDiNg oUtwARDs. LAteRAl siDes of pRoNotAl Disc sUbpARAllel iN bAsAl hAlf; wiDest iN ApicAl thiRD; withoUt ApophYseAl DepRessioNs. LAteRAl emARgiNAtioN of pRoNotUm NARRow AND complete; bAsAl emARgiNAtioN AbseNt; ANteRioR wiDelY iNteRRUpteD miDDle. BAse of pRoNotUm bisiNUAte. MiDDle of pRosteRNUm, iN mAles, coveReD bY pRomiNeNt setAe. PRoNotAl hYpomeRoN shiNY, with coARse pUNctURes (1.0 DiAmeteRs ApARt); sometimes with tUbeRcles oN lAteRAl eDge. INteRcoxAl pRocess of pRosteRNUm DepResseD ApicAllY. PRocoxAe NARRowlY sepARAteD (pRocoxA: iNteRcoxAl pRocess wiDth RAtio cA. 5.0).

ElYtRAl stRiAe Not impResseD, visible As teN Rows of coNspicUoUs pUNctURes oN eAch elYtRoN (siNgle DiAmeteR ApARt). INteRvAls 1, 3, 5, 7, 9 iN foRm of shARplY RAiseD coNtiNUoUs costAe; iNteRvAl 8 composeD of eloNgAteD tUbeRcles; 2, 4, 6, 10 flAt AND DUll, coveReD with micRoscUlptURe. ElYtRAl bAse v-like; Not emARgiNAte. EpipleURoN DilAteD ApicAllY, with RoUNDish AlveolA At level of foURth veNtRite; slightlY wiDeR thAN NeighboRiNg elYtRAl iNteRvAl; sitUAteD veNtRAllY, iNvisible iN DoRsAl Aspect. ElYtRAl hUmeRUs DeNtAte, stRoNglY pRotRUDiNg lAteRAD. ScUtellUm RelAtivelY smAll; tRiANgUlAR. MetAthoRAcic wiNgs AbseNt.

MesoveNtRite withoUt pRocesses. MetAveNtRite RelAtivelY shoRt (metAcoxAl cAvitY: metAveNtRite (betweeN iNseRtioNs of meso- AND metAcoxAe) leNgth RAtio cA. 2.4). PRocess of fiRst AbDomiNAl veNtRite NARRow (DistANces betweeN mesocoxAe: metAcoxAe RAtio cA. 0.7). Fifth AbDomiNAl veNtRite UNboRDeReD; coveReD bY fiNe pUNctURes (siNgle DiAmeteR ApARt).

Meso- AND metAtRochANteRs NeARlY eNtiRelY coveReD bY Respective femoRA; eqUippeD with A siNgle setAe. MetAcoxA RoUNDeD (wiDth: leNgth RAtio cA. 1.1). TARsi of both sexes NARRow. MAle pRotibiA with meDiAN cAvitY. UNDeRsiDe of mAle metAtibiA AND metAfemoRA coveReD with A Row of setAe. FemAle legs simple (pRotibiAe eveNlY wiDeNeD towARD Apex).

AeDeAgAl tegmeN wiDest meDiAllY; UNipARtite. PARAmeRes DiviDeD (Up to 0.3 of whole tegmeN leNgth). ClAvAe stRAight. PeNis wiDest meDiAllY; eveNlY NARRowiNg towARDs Apex NeAR Apex. BAsAl ApophYses of peNis RelAtivelY shoRt (cA. 0.25 of whole tegmeN leNgth).

MoRphologY of ovipositoR (boDY: ovipositoR leNgth RAtio cA. 7.0) AND femAle geNitAl tUbes coNsisteNt with those ReveAleD foR T. (T.) glandipleurum ( Fig. 5A, C, D ).

Distribution. This species hAs beeN collecteD iN the followiNg ecoRegioNs of SoUth AfRicA (MoUNtAiN ZoNe): DRAkeNsbeRg moNtANe gRAsslANDs, wooDlANDs AND foRests ( Fig. 7 ).

Figures

FIGURE 4. Habitus images of the studied species. Tragardhus (Tragardhus) biaplicalis Koch, 1956, holotype (A), T. (T.) glandipleurum Koch, 1956, syntype (B), T. (T.) jani sp. nov., holotype (C, F), T. (T.) majae sp. nov., paratype (D) and T. (T.) stigmaticus Koch, 1956, syntype (E, G).

FIGURE 5. Male and female terminalia of Tragardhus and Pseudemmallus. Ovipositor (A – D), genital tubes (C, D), aedeagal tegmen (E, G) and penis (F). Tragardhus (Tragardhus) glandipleurum Koch, 1956 (A, C – F), T. (Mitragardhus) nodosus (B) and Pseudemmallus aspericollis (G). Abbreviations: C 1 – C 3 — subsequent lobes of coxites, val—valvifer, ov—oviduct, bcbursa copulatrix, sp—spermatheca, par—paraproct, p—penis, cl—clavae.

FIGURE 7. Distribution of the genus Tragardhus.