Ummidia nidulans (Fabricius, 1787)

Godwin, Rebecca L. & Bond, Jason E., 2021, Taxonomic revision of the New World members of the trapdoor spider genus Ummidia Thorell (Araneae, Mygalomorphae, Halonoproctidae), ZooKeys 1027, pp. 1-165 : 1

Ummidia nidulans (Fabricius, 1787) Fig. 73 , Map 6

Aranea nidulans Fabricius, 1787: 343; female NEOTYPE designated herein (UMM297, deposited in AMNH) from Munro College, St. Elizabeth, Jamaica, 17.9244 -77.6869 4, coll. R. Physick v.1959; original holotype presumably lost.

Mygale nidulans Latreille, 1804: 166.

Cteniza nidulans Sells, 1837: 207.

Cteniza venatoria Koch, 1838: 12.

Actinopus nidulans Westwood, 1840: 173.

Actinopus venatorius Koch, 1850: 75.

Pachylomerus nidulans Ausserer, 1871: 147.

Pachylomerus nidulans Simon, 1892: 87.

Ummidia nidulans Rudloff, 1996: 48.

Diagnosis.

Ummidia nidulans is a relatively large species (CL 11-13.5), which can be differentiated from U. tibacuy , U. asperula , and U. neblina by the presence of a defined comb of long spinules on the retrolateral face of tarsus IV. Females can be further distinguished from all geographically proximate species by having a large number of distal endite cuspules (> 90 vs 30-54) and spermathecae which are straight with rounded, anteriorly facing bulbs.

Female neotype.

Specimen preparation and condition. Specimen preserved in 80% EtOH. Spermathecae removed, cleared, in vial with specimen. General coloration. Carapace, chelicerae, and legs dark reddish brown 2.5YR 2.5/4. Abdomen black 5YR 2.5/1, spinnerets dark brown 7.5YR 3/4. Cephalothorax. Carapace 13.48 long, 12.72 wide. Pars cephalica 9.26 long. Foveal groove procurved, 1.42 long, 3.29 wide. Eye tubercle moderate under ME. AER procurved. PER slightly recurved. Eye group 1.3 long, 2.89 wide, AME 0.6, PME 0.44, ALE 0.68, PLE 0.58. Sternum sparsely setose, thicker anteriorly, STRl 7.9, STRw 8.3. Chelicerae with anterior row comprising 12 teeth, posterior margin with nine teeth. Palpal endites with 24 cuspules spread across proximal half of endite and 103 smaller cuspules distally, ENDw 3.28, ENDl 5.32. Labium with 14 cuspules, LBw 2.77, LBl 1.86. Rastellum with many strong spines on process, none along margin, with small spines above process. Abdomen. Evenly setose with pale speckles forming patches at apodemes, lighter dorsally. Legs. F1 7.41; F1w 2.81; P1 5.11; Ti1 4.57; Mt1 4.47; Tr1 2.42; F3 6.8; F3w 3.47; P3 4.61; Ti3 4.16, Sd3 2.6; Mt3 2.72; Tr3 2.26; F4 7.83; F4w 3.44; P4 4.57; Ti4 4.57; Mt4 4.47; Tr4 2.42. Retrolateral face tarsus IV with defined comb of long spinules over length of tarsus. Pedipalps. PF 6.57, PP 4.14, PTi 4.44, PTr 3.68. Spermathecae simple on relatively long, weak stalks, bulbs facing anteriorly.

Variation, females.

(n = 5) CL 4.95-13.48, 11.14 ± 1.59; CW 4.65-12.72, 10.43 ± 1.5; Cap 3.75-10.11, 8.02 ± 1.12; ENDl 0.68-1.39, 1.16 ± 0.13; ENDw 1.15-2.89, 2.28 ± 0.31; STRl 2.98-8.12, 6.7 ± 0.96; STRw 2.98-8.3, 6.02 ± 1.18; LBl 1.07-2.19, 1.83 ± 0.2; LBw 1.22-3.11, 2.49 ± 0.34; F1 3.23-8.04, 6.58 ± 0.88; F1w 1.28-3, 2.5 ± 0.31; P1 2.05-5.25, 4.32 ± 0.59; Ti1 1.91-5.41, 4.28 ± 0.62; Mt1 1.18-3.75, 2.89 ± 0.45; Tr1 0.92-1.96, 1.61 ± 0.19; F3 2.75-7, 5.8 ± 0.78; F3w 1.58-3.65, 3.1 ± 0.39; P3 1.87-4.74, 3.9 ± 0.53; Ti3 1.55-4.21, 3.42 ± 0.5; Mt3 1.13-3.14, 2.46 ± 0.35; Tr3 1.12-3.13, 2.28 ± 0.34; F4 3.53-8.34, 7.03 ± 0.89; F4w 1.57-3.65, 3.01 ± 0.38; P4 1.94-4.78, 3.99 ± 0.53; Ti4 2.04-5.02, 4.1 ± 0.55; Mt4 1.81-4.57, 3.79 ± 0.53; Tr4 1.21-3.01, 2.29 ± 0.31; PF 2.88-7.44, 5.91 ± 0.8; PP 1.71-4.24, 3.58 ± 0.48; PTi 1.69-4.72, 3.85 ± 0.56; PTr 1.59-4.28, 3.36 ± 0.47.

Males.

Unknown.

Material examined.

Jamaica: 18.1192 -77.2977 9, 311 m a.s.l. (UMM0476, 13.viii.1923, 1♀, JT Lloyd, AMNH); Clarendon: Mosley Hall , 18.108 -77.2986 9, 309 m a.s.l. (UMM0305, i.1966, 1♀, R Kern, AMNH); Portland: Cinchona , 18.1056 -76.68 6, 1553 m a.s.l. (UMM0201, 1♀, CT Brues, MCZ); St. Elizabeth: Munro College , 17.9244 -77.6869 4, 774 m a.s.l. (UMM0297, 1-31.v.1959, 1♀, R Physick, AMNH); AMNH).

Figures

Map 6. Locality records for South American and Caribbean species.

Figure 73. Ummidia nidulans (Fabricius, 1787) from St. Elizabeth, Jamaica. Female neotype (UMM 297) A female habitus illustration B cleared spermathecae. Scale bars: 4.0 mm (A), 1.0 mm (B).