Vexillum szobbiensis ( Halaváts, 1884 )

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 54

Vexillum szobbiensis ( Halaváts, 1884)

Figs 14C 1 –C 2 , D

* Mitra (Costellaria) Szobbiensis Halaváts 1884: 178 , pl. 4. fig. 5.

Vexillum (Costellaria) szobbiensis Halaváts — Csepreghy-Meznerics 1956: 413, pl. 9, figs 13–14.

Mitra (Vexillum) intermittens szobbiensis Halaváts, 1884 — Strausz 1966: 368, pl. 25, figs 6–8.

Mitra (Vexillum) intermittens Hoernes & Auinger, 1880 — Strausz 1966: 368, pl. 25, figs 9–10 [non Vexillum intermittens ( Hoernes & Auinger, 1880) ].

Type material. Lectotype (designated herein): Illustrated in Halaváts (1884, pl. 4. fig. 5), Inv. nr. M107, Mining and Geological Survey of Hungary, SL: 16.9 mm, MD: 5.0, Szob ( Hungary), figs 14C 1 –C 2 . Paralectotype: Inv. nr. M107, Mining and Geological Survey of Hungary, SL: 14.0 mm, MD: 4.2 mm, Szob ( Hungary), fig. 14D .

Revised description. Shell, moderately large, slender fusiform, with high spire. Protoconch unknown. Teleoconch of nine whorls. Early spire whorls weakly convex, with periphery slightly below mid-whorl, passing to subcylindrical on late spire whorls, separated by weakly incised suture. Early teleoconch whorls nearly smooth, with indistinct opisthocline axial swellings, abapically strengthening into wide-spaced, low, convex axial ribs on sixth to eighth teleoconch whorls. Onset of spiral sculpture coinciding with axial sculpture, consisting of numerous weak spiral cords, most prominent in axial interspaces, but also overriding axial ribs. Last whorl short, subcylindrical, moderately constricted. Aperture wide. Columellar callus indistinct. Columella with four oblique columellar folds. Siphonal canal, moderately long, wide, straight, with shallow siphonal notch.

Shell measurements and ratios. SL: 14.0– 16.9 mm, MD: 4.2–5.0 mm; AA: 25–28°, SL/MD: 3.5, AL/AW: 3.9, AH/S: 2.2.

Discussion. This species is unique amongst the Paratethyan costellariids in being characterised by a change of sculpture from nearly smooth on early spire whorls to distinct axial and spiral sculpture on later whorls, with the transition occurring on about the seventh teleoconch whorl.

Palaeoenvironment. The mollusc fauna and lithofacies of Szob suggest a sandy coastal environment with sea grass ( Dulai 1996).

Distribution in Central Paratethys. Badenian (middle Miocene): Pannonian Basin: Szob ( Hungary) ( Strausz 1966).

Figures

FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 14 A1–A2. Pusia paraleucozona (Boettger, 1906), Drnovice u Vyškova (Czech Republic), NHMW 1865/0036/0010. B1–B2. Pusia paraleucozona (Boettger, 1906), lectotype, Coşteiu de Sus (Romania), SMF 360345. C1–C2. Pusia szobbiensis (Halaváts, 1884), lectotype, Inv. nr. M107, Szob (Hungary). D. Vexillum szobbiensis (Halaváts, 1884), paralectotype, Inv. nr. M107, Szob (Hungary). E1–E2. Vexillum pseudoschafferi Biskupič, 2020, holotype, Z 40033, Rohožník–Konopiská (Slovakia). E1–E2. Vexillum pseudoschafferi Biskupič, 2020, paratype, Z 40034, Rohožník–Konopiská (Slovakia). G. Vexillum pseudoschafferi Biskupič, 2020, paratype, Z 40035, Rohožník–Konopiská (Slovakia).