Bembidion tetracolum tetracolum Say, 1823

Bousquet, Yves, 2012, Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico, ZooKeys 245, pp. 1-1722 : 372-373

Bembidion tetracolum tetracolum Say, 1823

Carabus littoralis Olivier, 1795: [35] 110 [secondary homonym of Bembidion litoralis (Olivier, 1790)]. Type locality: "environs de Paris, sur les bords de la Seine [France]" (original citation). Syntype(s) location unknown (possibly in MHNP). Note. The use of a single or double consonant in the spellings of species-group names is deemed to be identical (ICZN 1999: Article 58.7); therefore the two Olivier’s names are homonyms.

Bembidium tetracolum Say, 1823a: 89. Type locality: "Arlington [Middlesex County], Mass[achusetts]" (neotype label). Neotype (♂), designated by Lindroth and Freitag (1969: 336), in MCZ [# 33068]. Synonymy established, under the name Bembidion ustulatum (Linnaeus) sensu auctorum (= Bembidion tetracolum Say), by Fassati (1950: 42).

Bembidion rupicola nactum Casey, 1918: 77. Type locality: "lower Hudson Valley, New York" (original citation). Lectotype (♀), designated by Lindroth (1975: 119), in USNM [# 36943]. Synonymy established by Lindroth (1954b: 126).

Distribution.

This Palaearctic subspecies is adventive in North America where it is known from Newfoundland (Lindroth 1955a: 62, as Bembidion ustulatum ) to western Montana (Russell 1968: 54), including southern Quebec and the Ontario Peninsula (Lindroth 1963b: 332), south to northern Utah (Salt Lake County, CMNH), west-central Nebraska (Keith County, Foster F. Purrington pers. comm. 2010), and central Virginia (Davidson 1995: 16); also in the Pacific Northwest from southwestern British Columbia (Lindroth 1963b: 332) to northern California (Notman 1929b: 222; Tehama County, Foster F. Purrington pers. comm. 2010). The records from Colorado (Wickham 1902: 233, as Bembidion ustulatum ), “Tennessee,” and "North Carolina" (Bousquet and Larochelle 1993: 138) need confirmation; that from “Texas” (Lindroth 1955a: 62) could be based on a mislabeled specimen. The first inventoried specimen collected on this continent was found prior to 1823, likely along the east coast. The species has been unearthed from a xvii Century colonial site in Boston, Massachusetts (Bain 1998: 39), suggesting that the species was established on this continent long before Thomas Say collected his specimens. The species is also adventive in southeastern Australia (Moore et al. 1987: 136).

Records.

FRA: PM CAN: BC, NB, NF, NS, ON, PE, QC USA: CA, CT, DC, IA, IL, IN, MA, ME, MI, MN, MT, ND, NE, NH, NJ, NY, OH, OR, PA, RI, SD, UT, VA, VT, WA, WI [CO, NC, TN, TX] - Adventive

Note.

This species has been known for a long time under the name Bembidion ustulatum (Linnaeus, 1758). Lindroth (1957b: 334-335) suggested to drop the name to promote stability. Four other subspecies of Bembidion tetracolum are found in the Palaearctic Region.