Calapnita bugis, Bernhard A. Huber, 2017

Bernhard A. Huber, 2017, Revision and cladistic analysis of the Southeast Asian leaf-dwelling spider genus Calapnita Simon (Araneae, Pholcidae), Zootaxa 4219 (1), pp. 1-63 : 40-42

Calapnita bugis sp. nov. Figs 180–187

“ Calapnita vermiformis ” (misidentification; only specimens from Sulawesi): Deeleman-Reinhold 1986a: 45; 1986b: 212. Huber 2011: 48, figs 141, 147–152.

Diagnosis. Distinguished from most species of vermiformis group (except C. dayak ) by wide obtuse tip of ventral flap of procursus ( Figs 180–181 ); from most (except C. saluang ) also by thin proximal part of bipartite cheliceral apophysis ( Fig. 184 ) and by strong and regular fringes at tip of embolus; from most (except C. bohol ) also by oval pore plates in transversal orientation ( Fig. 187 ); from several species also by narrow and distinct epigynal ‘knob’ ( Fig. 186 ).

Etymology. Named for the Buginese people, an ethnic group in southwestern Sulawesi; noun in apposition.

Material examined. Holotype. INDONESIA-SULAWESI: ♂, RMNH ( ARA 17817), South Sulawesi, 55 km from Udjung Pandang, 23 km from Camba [~ 5.05°S, 119.74°E], under leaves in karst forest, 9 & 13–14.viii.1980 (C.L. & P.R. Deeleman).

Other material. INDONESIA-SULAWESI: 3♂ 3♀, RMNH ( ARA 17420), same data as holotype ; 1♂ 1♀, RMNH ( ARA 17426), same data but only 9.viii.1980 . 2♂ 3♀, RMNH ( ARA 17423), North Sulawesi, Dumoga, near Doloduo [~ 0.517°N, 123.917°E], primary forest, from leaves, 27–30.viii.1982 (collector not given).

Assigned tentatively. INDONESIA-SULAWESI: 2♂ 2♀, RMNH ( ARA 17425), Central Sulawesi, Palu, Marena, Lore Lindu Reserve [~ 1.3°N, 120, 083°E], “dark little stream valley”, from leaves, 24.vii.1982 (collector not given).

Description. Male (holotype)

MEASUREMENTS. Total body length 5.2, carapace width 0.65. Leg 1 missing, tibia 2: 5.5, tibia 3: 3.4, tibia 4: 6.0. Distance PME-PME 200 µm, diameter PME 80 µm, distance PME-ALE ~30 µm; no trace of AME.

COLOR. Entire animal very pale gray to whitish, legs with brown patellae and tibia-metatarsus joints.

BODY. Habitus as in close relatives (cf. Figs 120, 122 ); ocular area barely elevated, each triad on very low hump; carapace without median furrow; clypeus unmodified; sternum barely wider than long (0.48/0.46), unmodified.

CHELICERAE. As in Figs 184–185 , apophyses near lamellae clearly bipartite, proximal parts slender (similar to C. saluang ), with pair of barely visible lateral processes; without modified hairs; without stridulatory ridges.

PALPS. In general similar to C. vermiformis (cf. figs 139 and 140 in Huber 2011); trochanter apophysis slender, as in Fig. 182 ; femur as in Fig. 183 , distal process at 44% of femur length; tibia length/diameter 0.51/0.25; procursus as in Figs 180–181 , with simple ventro-distal flat sclerite; bulb length 0.40; embolus length 0.60; embolus tip with strong and regular semi-transparent fringes.

LEGS. Without spines and curved hairs; few vertical hairs.

Male (variation). Tibia 1 in three other males: 7.7, 7.7, 7.8. The specimens from Lore Lindu Reserve are assigned tentatively because the tip of the procursus differs slightly (ventro-distal flat sclerite directed slightly more towards ventral).

Female. In general similar to male; eye triads slightly closer together (distance PME-PME 180 µm). Tibia 1 in 3 females: 6.3, 6.5, 6.7. Epigynum as in Fig. 186 , weakly sclerotized triangular plate with small and slender anterior ‘knob’; internal genitalia as in Fig. 187 , much longer than wide, with small elongated pore plates in transversal orientation far from posterior margin.

Distribution. Apparently widespread in Sulawesi ( Fig. 283 ).

Figures

FIGURE 283. Known distribution of the Calapnita vermiformis group, part 1 (cf. Fig. 284). The Java record of C. lawangan is based on specimens assigned tentatively.

FIGURES 172 – 187. Calapnita lawangan sp. nov. (172 – 179; RMNH, ARA 17815 – 16) and C. bugis sp. nov. (180 – 187; RMNH, ARA 17420). 172 – 173, 180 – 181. Left procursi, prolateral and retrolateral views (arrow points at distinctive undulating ridge). 174, 182. Left palpal trochanters, retrolateral views. 175, 183. Left palpal femora, prolateral views. 176 – 177, 184 – 185. Right male chelicerae, frontal and lateral views. 178, 186. Epigyna, ventral views. 179, 187. Cleared female genitalia, dorsal views. Scale lines: 0.3 mm.

FIGURES 120 – 127. Live specimens, Calapnita vermiformis group. 120 – 121. C. bario sp. nov., ♂ and ♀ with egg-sac from Bario. 122 – 123. C. magaseng sp. nov., ♂ and ♀ with egg-sac from Mount Penrissen. 124 – 125. C. saluang Huber, ♂ from Berastagi and ♀ with egg-sac from Phanom Bencha. 126 – 127. C. vermiformis Simon, ♂♂ from Mount Isarog and Mount Banahaw.