Cardiacephala modesta ( Cresson, 1930 ) Ferro & Marshall, 2018

Ferro, Gustavo Borges & Marshall, Stephen A., 2018, A revision of the Neotropical ant-like genus Cardiacephala Macquart, including Plocoscelus Enderlein syn. nov. (Diptera: Micropezidae, Taeniapterinae), Zootaxa 4429 (3), pp. 401-458 : 432-434

Cardiacephala modesta ( Cresson, 1930) new combination

( Figs. 50–52 )

Plocoscelus modestus Cresson, 1930: 334 .

Description: Body length 7 mm, wing 4 mm. Palpus mostly brown, white on outer face. Clypeus brown, shiny. First flagellomere brown and with golden microsetulae. Fronto-orbital plates brown. Postpronotal lobe yellow, shiny. Row of katepisternal setae brownish. Fore tibia dark brown. Hind femur narrowly white basally. Fore tarsus dark brown with gold/whitish pubescence ventrally at base of tarsomere one. Oviscape yellow and shiny, apical portion darkened. Epandrium yellow.

Head: Palpus broadened basally and gradually tapering to apex. Clypeus short, length less than half of width. Two frontal setae. Inner vertical setae inserted in strong tubercles. Paracephalon smooth, without swelling (not extending beyond the vertex). Vertex rounded, irregular due to the strong tubercles on epicephalon. Supracervical line of setae black.

Thorax: Mesonotum with golden pruinosity. Scutellum upturned, triangular. Femora with sparse black setulae. Mid and hind tibia flattened, but sulcus on outer face weak. Wing stigmatal and discal band delimited, incomplete apical pigmentation with three darkened spots. Crossvein r-m with small dark spot. R4+5 and M1+2 meeting at the wing margin. A1+CuA2 as long as CuA2. Female cervical sclerite smooth, without swelling.

Abdomen: Tergites with sparse golden setulae. Female abdomen: Single spermathecal duct (slightly damaged) uniformly thick, ending in a cylindrical (peanut-like) spermatheca. Paired spermathecal duct divided into a narrow basal half and a broader distal half. Paired spermathecal duct stems thin and smooth basally, followed by a small rounded swelling and a striate duct with a small appendix ( Fig. 51 ). Paired spermathecae spherical, somewhat flattened basally and deeply invaginated from apex to base. Male abdomen: Ejaculatory apodeme somewhat thin and about same length as epandrium, blade narrowed, about as wide as sperm pump. Phallapodeme extending to anterior limit of hypandrium. Distiphallus long, longer than epandrium, ending in a phallic bulb, phallic subequal in length and width ( Fig. 52 ).

Type material: Holotype (1 ♀, ANSP, only images examined): GUATEMALA. Puerto Barrios, 20. IV.1924 (white label) Type No. 6388, E.T. Cresson Jr. (red label).

Material examined: COLOMBIA. Amazonas, PNN Amacayacu Mocagua, 3º23’1” N 70º06’1” W, 300 m, M. 853, 14–21.VIII.2000, A. Parente (1 ♀, IAVH); Chocó, Ensenada de Utria Natl. Pk., Visitor Centre, 0–10 m, Malaise trap, 5–19.VII.2000, J. Pérez (9 ♀, DEBU 00138532, DEBU 00138533, DEBU 00138535, DEBU 00138536, DEBU 00138537, DEBU 00138538, DEBU 00138539, DEBU 00138540, DEBU 00138541, IAVH); Chocó, PNN Utriá, Centro de Visitantes, 6º01’09” N 77º20’55” W, 2 m, 15–30.XII.2000, M. 333, J. Pérez (7 ♀, IAVH); Chocó, PNN Utriá, Centro de Visitantes, 6º01’09” N 77º20’55” W, 2 m, 28.IX–15.X.2000, M. 815, J. Pérez (9 ♀, 1 ♂, IAVH); Chocó, PNN Utriá, Centro de Visitantes, 6º01’09” N 77º20’55” W, 2 m, 15–30.X.2000, M. 816, J. Pérez (14 ♀, 3 ♂, IAVH); Chocó, PNN Utriá, Centro de Visitantes, 6º01’09” N 77º20’55” W, 2 m, 15– 30.X.2000, M. 816, J. Pérez (1 ♀, DEBU 01086318, IAVH); Chocó, PNN Utriá, Centro de Visitantes, 6º01’09” N 77º20’55” W, 2 m, 2–15.VIII.2000, M. 818, J. Pérez (8 ♀, IAVH); Chocó, PNN Utriá, Centro de Visitantes, 6º01’09” N 77º20’55” W, 2 m, 15.VIII–7.IX.2000, M. 820, J. Pérez (7 ♀, 1 ♂, IAVH); Chocó, PNN Utriá, Centro de Visitantes, 6º01’09” N 77º20’55” W, 2 m, 7–28.IX.2000, M. 823, J. Pérez (9 ♀, 2 ♂, IAVH); Chocó, PNN Utriá, Centro de Visitantes, 6º01’09” N 77º20’ W, 2 m, 1–15.XI.2000, M. 1341, J. Pérez (4 ♀, IAVH); Chocó, PNN Utriá, Centro de Visitantes, 6º01’09” N 77º20’ W, 2 m, 26.XII.2000 – 1.II.2001, M. 1343, J. Pérez (9 ♀, 2 ♂, IAVH); Chocó, PNN Utriá, Centro de Visitantes, 6º01’09” N 77º20’ W, 2 m, 26.XII.2000 – 1.II.2001, M. 1343, J. Pérez (1 ♀, IAVH, sequence DEBUA 281–17); Chocó, PNN Utriá, Centro de Visitantes, 6º01’09” N 77º20’55” W, 2 m,

26.XII.2000 – 1.II.2001, M. 1465, J. Pérez (10 ♀, 2 ♂, IAVH); Chocó, PNN Utriá, Cocalito, 6º01’09” N 77º20’55” W, 20 m, 7–28.IX.2000, J. Pérez (1 f #, IAVH).

Distribution: Colombia, Guatemala.

Figures

FIGURES 50–52. Cardiacephala modesta. 50, female, dorsolateral view; 51, female genitalia, ventral view; 52, male sternite 5 and genitalia, ventral and lateral view, respectively.