Crusopimpla tadushiensis Kopylov, Spasojevic & Klopfstein, 2018

Kopylov, D. S., Spasojevic, T. & Klopfstein, S., 2018, New ichneumonids (Hymenoptera, Ichneumonidae) from the Eocene Tadushi Formation, Russian Far East, Zootaxa 4442 (2), pp. 319-330 : 323-325

Crusopimpla tadushiensis Kopylov, Spasojevic & Klopfstein sp. nov.

Figure 3

Holotype. PIN, 3364/277, part with counterpart. Imago, female in dorsal aspect. Head absent, meso- and metasoma well preserved, fore wings almost complete, hind wings absent, legs incomplete, ovipositor sheath complete. Yushangou, Russian Far East, Eocene, Tadushi Formation.

Additional material. Paratype, PIN, 3364/41, part with counterpart, isolated fore wing, poorly preserved. Ugolny, Russian Far East, Eocene, Tadushi Formation.

Etymology. This species is named after the type locality, the Tadushi Formation.

Description. Coloration. Body, legs, wing veins, ovipositor uniformly dark.

Mesosoma. Mesonotum 0.8× as long as wide, 2.8× as long as scutellum. Notauli weak, converging, reaching the middle of mesonotum. Propodeum with median longitudinal carinae complete, divided in areas by basal and apical transverse carinae (apical transverse carina very weak, so border between areola and petiolar area somewhat unclear). Areola hexagonal, as wide as long; basal area trapezoid, 2.0× as wide as long and 0.6× as wide as areola; petiolar area hexagonal, 1.8× as wide as long and 2.3× as wide as areola. Lateral longitudinal carina joins median one at mid-length of petiolar area; probably three lateral areas developed. Ventrolateral aspect of propodeum not preserved, pleural areas thus not clear.

Wings. Fore wing 2.9× as long as wide; pterostigma 2.5× as long as wide; 1-Rs&1-M slightly arched; 1mcu&2-Rs+M curved, with no sharp bend or traces of 1-Rs+M (ramulus); r-rs issues at mid-length of pterostigma, straight, 1.6× pterostigma width; areolet quadrate, 2.2× as long as wide, with 2+3-M 1.8× 2-Rs, 1.2× r-m, 5.3× 4- M; 1cu-a interstitial.

Legs. Hind femur approximately 2.5× as long as wide, 1.7× as wide as hind tibia, which is ~5.0× as long as wide. First hind tarsomere 0.65× as long as tibia and 2.4× as long as second tarsomere.

Metasoma. Metasoma fusiform, 2.2× as long as wide and 1.8× as long as mesosoma. T1 (first metasomal tergite) short, 0.9× as long as wide, comprises 0.2 of metasoma length; with strong dorsal carinae converging on basal half, remainder parallel and as far apart as either is from lateral margin of T1. Following tergites strongly transverse, of nearly equal lengths, with no apparent sculpture (which might be a preservation artefact). Ovipositor sheath rigid, stout, with transverse carinulae, 0.26× as long as fore wing.

Measurements. Mesosoma length 3.0 mm, metasoma length 5.5 mm, ovipositor length (sheath) 2.1 mm. Fore wing length 7.8 mm, width 2.7 mm.

Figures

FIGURE 3. Crusopimpla tadushiensis sp. nov.: a–c—holotype PIN, 3364/277: a—part; b—our interpretation of the fossil, along with the wing vein (left), wing cell (right) nomenclature and the structure of mesosoma: Sct 2 —mesoscutum, Scl 2 —mesoscutellum, N 2 ax—axillary trough of mesonotum, n—notauli, N 3 —metanotum (postscutellum), N 3 ax—axillary trough of metanotum, ba—basal area, a—areola, pa—petiolar area, la 1 —1-st lateral area, la 2 —2-nd lateral area, la 3 —3-rd lateral area; c—SEM photo of propodeum; d—paratype PIN, 3364/41 (d).