Dendrocellus crassus (Andrewes) Liang & Kavanaugh, 2007

Liang, Hongbin & Kavanaugh, David H., 2007, Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species, The Coleopterists Bulletin (Oxford, England) 61 (1), pp. 1-39 : 1-39

Dendrocellus crassus (Andrewes) , new combination

( Figs. 5 , 45 , 48, 50 , 53, 56 , 83–84 )

Desera crassa Andrewes 1931: 477 (Borneo 5 Pulau Kalimantan)

Specimens examined. Total 18 specimens: Indonesia : 1 female ( ISNB), ‘‘ Coll. I. R. Sc. N. B. Borneo 1900, Pontianak, ex coll. Oberthur’ ’; 1 female ( ISNB), ‘‘ Coll .

18 I. R. Sc. N. B. Java – Togoe, 1902, ex. Coll. Oberthur’’; 1 female (ISNB), ‘‘Beokit Gabah, Z. W. K. Sumatra, leg. H. Lucht, I.1919 ’’/‘‘Coll. I. R. Sc. N. B. Sumatra’’; 1 male (ZMHU), ‘‘W. Sumatra, No. 18, VII.1991 ’’. Malaysia: 2 females (ZMHU), ‘‘Near Keningau, Is. Borneo, Sabah, E. Malaysia, 16 II 1989, M. Itoh’’/‘‘ Desera geniculata (Klug) det. N. Ito, 1993’’; 1 male and 1 female (ISNB), ‘‘ Kina Balu , Borneo’’/‘‘Collection P. Dupuis’’/‘‘Coll. R. I. Sc. N. B. Borneo’’; 1 male and 1 female (ISNB), ‘‘ Kina balu , Borneo 1,500 m, H. Hille, Berlin SW11’’/ ‘‘R. I. Sc. Nat. Belg. I.G. 16.500’’/‘‘Coll. R. I. Sc. N. B. Borneo’’; 1 female (ISNB), ‘‘Coll. I. R. Sc. N. B. Nord Borneo, Kina Balu, Fr. Buffat, 1904 ’’; 1 female (ISNB), ‘‘Coll. I. R. Sc. N. B. Indonesie, N. Borneo, Mont Kina Balu , 5.VIII.1903, leg. John Waterstradt, ex coll. Oberthur IG. 18.293’’. Kalimantan (Malaysia or Indonesia): 1 male (ISNB), ‘‘Borneo’’/‘‘Collection P. Dupuis’’/‘‘Coll. R. I. Sc. N. B. Borneo’’; 1 female (ISNB), ‘‘Borneo’’/‘‘Collection P. Dupuis’’/ ‘‘ Desera unidentata M.L. det. P. Dupuis, 1913’’/‘‘ Desera unidentata ’’/‘‘Coll. R. I. Sc. N. B. Borneo’’; 1 male (ISNB), ‘‘Borneo’’/‘‘Reg. Mus. Hist. Nat., Belg. I.G. 10294’’/‘‘ longicollis Dj’’/‘‘G. Fagel’’/‘‘G. Fagel det. Desera longicollis Dej’’/‘‘Coll. R. I. Sc. N. B. Borneo’’; 1 male and 2 females (CAS), ‘‘Borneo, M. L. Hauschild’’/ ‘‘Van Dyke collection’’/‘‘ Dendrocellus crassus Andrewes 1931 , det. Liang H. B. 2005.3’’.

Description. Andrewes (1931) provided a detailed description and illustration. We simply add the following: length 9.4–10.4 mm, width 2.9–3.3 mm. Apical (not basal as stated in original description) four fifths of Ant 1 black; femora brown, with apical sixth or seventh black; Ant 1/Ant 3 5 3.45 (3.33–3.56); HW/PW 5 1.17 (1.12–1.21); PL/PW 5 1.53 (1.48–1.57); EW/PW 5 2.36 (2.18–2.42); EL/EW 5 1.72 (1.64–1.81); labrum with medial lobe slightly protruded anteriorly; genae short, slightly tumid; eyes moderately large, 2.5 times as long as genae; elytral punctation as in Fig. 45 ; protarsomere 3 symmetrical in male ( Fig. 53 ); tarsal claws with 5–6 pectinations, length of the longest pectination subequal to the width of claw base (similar to Fig. 63 ); last sternum as in Fig. 48 or 50. Aedeagus with apical lamella thin, linguiform ( Figs. 83–84 ).

Distribution. Indonesia (Sumatra), Malaysia (N. Kalimantan).

Remarks. We have not examined the type (in BMNH). Members of this species are readily recognized by their elytral intervals sparsely punctate ( Fig. 45 ), femora with apical sixth black, and tibiae completely black.

Figures

Figs. 1–12. Habitus of Dendrocellus species. 1) D. australis (male, Congo); 2) D. coelestinus (syntype); 3) D. coelestinus (type of D. paralellus); 4) D. confusus (male, Thailand); 5) D. crassus (male, Borneo); 6) D. elegans (male, New Guinea); 7) D. rugicollis (lectotype); 8) D. geniculatus (holotype); 9) D. geniculatus (holotype of D. gilsoni); 10) D. gestroi (lectotype); 11) D. inexpectus (holotype); 12) D. javanus (holotype). Scale line 5 2.0 mm.

Figs. 44–46. Elytral punctation of Dendrocellus species. 44) D. geniculatus (male, Tibet);

Figs. 47–50. Apical margin of terminal sternum of Dendrocellus species. 47) and 49) D. geniculatus (Tibet); 48) and 50) D. crassus (Borneo); 47 and 48 5 males; 49 and 50 5 females. Scale line 5 1.0 mm.

Figs. 51–56. Tarsomeres 1–4 of right protarsus of Dendrocellus species (dorsal view). 51) and 54) D. viridipennis (Congo); 52) and 55) D. geniculatus (Tibet); 53) and 56) D. crassus (Borneo); 51 to 53 5 males; 54 to 56 5 females. Scale line 5 1.0 mm.

Figs. 73–90. Male genitalia of Dendrocellus species. 73) and 74) D. confusus (Thailand); 75) and 76) D. rugicollis (lectotype); 77) and 78) D. coelestinus (Thailand); 79) and 80) D. nepalensis (Pedong, India); 81) and 82) D. kulti (Hubei, China); 83) and 84) D. crassus (Borneo); 85) and 86) D. nigripennis (holotype); 87) and 88) D. geniculatus (Pedong, India); 89) and 90) D. javanus (holotype). Scale lines 5 1.0 mm, scale line A for Figs. 73, 75, 77, 79, 81, 83, 85, 87, 89 (apical part, dorsal view); scale line B for Figs. 74, 76, 78, 80, 82, 84, 86, 88, 90 (left lateral view).

Figs. 57–72. Tarsal claws of hind tarsi of Dendrocellus species and Drypta species. 57) D. rugicollis (lectotype); 58) D. elegans (male, New Guinea); 59) D. schultzei (lectotype); 60) D. sinicus (holotype); 61) D. inexpectus (holotype); 62) D. javanus (male, Java Occid., paratype); 63) D. nigripennis (holotype); 64) D. geniculatus (male, Tibet); 65) D. australis (male, Congo); 66) D. queenslandicus (holotype); 67) D. confusus (male, Thailand); 68) D. nepalensis (male, Pedong, India); 69) D. coelestinus (syntype); 70) D. micropectinatus (male paratype, Congo); 71) Drypta longicollis Macleay 1825 (male, Thailand); 72) Drypta lineola virgata Chaudoir 1850 (male, Vietnam). Scale line 5 0.1 mm.