Dendrocellus javanus Liang and Kavanaugh, 2007

Liang, Hongbin & Kavanaugh, David H., 2007, Review of the Genus Dendrocellus Schmidt-Göbel (Coleoptera: Carabidae: Dryptini), with Descriptions of Seven New Species, The Coleopterists Bulletin (Oxford, England) 61 (1), pp. 1-39 : 1-39

Dendrocellus javanus Liang and Kavanaugh , new species

( Figs. 12 , 34 , 62 , 89–90 )

Types. Holotype: a male ( ISNB), ‘‘ Java, Seokaboemi’’/‘‘ R. I. Sc. N. B. 19.458, Coll. Le Moult’ ’/‘‘Coll. R. I. Sc. N. B. Java’ ’/’’ HOLOTYPE, Dendrocellus javanus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006’’ [red label]. Paratypes (total 5 specimens): 1 female ( ISNB), ‘‘ Java, Soc. Ent. Belg., Coll. Putzeys’ ’/‘‘Coll. R. I. Sc. N. B. Java’ ’/’’ PARATYPE, Dendrocellus javanus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006’’ [yellow label]; 1 male and 1 female ( ISNB), ‘‘ Java’ ’/‘‘ R. Mus. Hist. Nat. Belg. I.G. 12.595’’/‘‘ Coll . R. I. Sc. N. B. Java’ ’/’’ PARATYPE, Dendrocellus javanus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006’’ [yellow label]; 1 female ( ISNB), ‘‘ Java’ ’/‘‘ Soc. Ent. Belg. Coll. Putzeys’ ’/‘‘ Coll . R. I. Sc. N. B. Java’ ’/’’ PARATYPE, Dendrocellus javanus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006’’ [yellow label]; 1 male ( ZMHU), ‘‘ Java Occid. Dr. Standgr. ’’/‘‘ Dendrocellus geniculatus Klug’ ’/’’ PARA- TYPE, Dendrocellus javanus Liang & Kavanaugh n. sp., designated by H. B. Liang & D. H. Kavanaugh 2006’’ [yellow label] .

Type locality. Java, Indonesia .

Diagnosis. Femora brown with apical fifth or sixth black; tarsal claws ( Fig. 62 ) with longest pectination subequal to width of tarsal claw base; elytra with outer angles slightly dentate; aedeagus with apical lamella moderately long, slightly digitiform, not thickened at apex ( Figs. 89–90 ).

Description. Length 9.2–10.2 mm, width 2.7–2.9 mm. Black, head, pronotum, and elytra with green luster, mixed with slight blue luster; apical third of Ant 1 and basal two thirds of Ant 3 black, basal two thirds of Ant 1, apical third of Ant 3, and Ant 4–11 brown or yellow; apical fifth or sixth of femora black, remainder brown or yellow; tibiae and tarsi brown or yellow. Head ( Fig. 34 ) wider than pronotum, HW/PW 5 1.18 (1.13–1.23); vertex convex, coarsely punctate; Ant 1/ Ant 3 5 3.35 (3.24–3.46); labrum with medial lobe slightly protruded anteriorly; genae moderately tumid; eyes large, about 2.2 times as long as genae. Pronotum densely punctate, narrow, PL/PW 5 1.46 (1.40–1.50); lateral margins gradually widened from apical angles to middle, strongly sinuate before basal angles; basal angles protruded laterally; basal foveae quite deep; middle line shallow. Elytra long, slightly widened from base to apical third, then narrowed to apex in both sexes (more so in female); EL/EW 5 1.82 (1.70–1.86), EW/PW 5 2.23 (2.12–2.32); outer angles slightly dentate (similar to Fig. 40 ); interval flat, densely punctate. Protarsomere 3 slightly asymmetrical in male; tarsal claw with 5–6 long pectinations, length of the longest pectination subequal to width of claw base ( Fig. 62 ). Aedeagus with apical lamella moderately long, slightly digitiform, unthickened at apex ( Figs. 89–90 ).

Distribution. Presently known only from type locality.

Etymology. The new species is named for Java, its type locality.

Remarks. Members of this new species are similar to those of D. geniculatus (Klug) in having tumid genae, green body luster, and similar femoral and tibial color, but differ in having the ratio of Ant 1/Ant 3 smaller and the lamella of aedeagus not thickened ( Fig. 90 ). They are also similar to adults of D. ternatensis Chaudoir in the form of male genitalia, but differ from the latter in femoral color and in having more tumid genae ( Fig. 34 ) and smaller ratio of Ant 1/Ant 3.

Figures

Figs. 1–12. Habitus of Dendrocellus species. 1) D. australis (male, Congo); 2) D. coelestinus (syntype); 3) D. coelestinus (type of D. paralellus); 4) D. confusus (male, Thailand); 5) D. crassus (male, Borneo); 6) D. elegans (male, New Guinea); 7) D. rugicollis (lectotype); 8) D. geniculatus (holotype); 9) D. geniculatus (holotype of D. gilsoni); 10) D. gestroi (lectotype); 11) D. inexpectus (holotype); 12) D. javanus (holotype). Scale line 5 2.0 mm.

Figs. 31–34. Head of Dendrocellus species. 31) D. queenslandicus (holotype); 32) D. ternatensis (male, Batchian, Indonesia); 33) D. rugicollis (lectotype, India); 34) D. javanus (holotype). Scale line 5 1.0 mm.

Figs. 57–72. Tarsal claws of hind tarsi of Dendrocellus species and Drypta species. 57) D. rugicollis (lectotype); 58) D. elegans (male, New Guinea); 59) D. schultzei (lectotype); 60) D. sinicus (holotype); 61) D. inexpectus (holotype); 62) D. javanus (male, Java Occid., paratype); 63) D. nigripennis (holotype); 64) D. geniculatus (male, Tibet); 65) D. australis (male, Congo); 66) D. queenslandicus (holotype); 67) D. confusus (male, Thailand); 68) D. nepalensis (male, Pedong, India); 69) D. coelestinus (syntype); 70) D. micropectinatus (male paratype, Congo); 71) Drypta longicollis Macleay 1825 (male, Thailand); 72) Drypta lineola virgata Chaudoir 1850 (male, Vietnam). Scale line 5 0.1 mm.

Figs. 73–90. Male genitalia of Dendrocellus species. 73) and 74) D. confusus (Thailand); 75) and 76) D. rugicollis (lectotype); 77) and 78) D. coelestinus (Thailand); 79) and 80) D. nepalensis (Pedong, India); 81) and 82) D. kulti (Hubei, China); 83) and 84) D. crassus (Borneo); 85) and 86) D. nigripennis (holotype); 87) and 88) D. geniculatus (Pedong, India); 89) and 90) D. javanus (holotype). Scale lines 5 1.0 mm, scale line A for Figs. 73, 75, 77, 79, 81, 83, 85, 87, 89 (apical part, dorsal view); scale line B for Figs. 74, 76, 78, 80, 82, 84, 86, 88, 90 (left lateral view).

Figs. 35–43. Left elytron apex of Dendrocellus species. 35) D. kulti (male, Yunnan); 36) D. nepalensis (male, Pedong, India); 37) D. rugicollis (lectotype, India); 38) D. micropectinatus (holotype); 39) D. australis (male, Congo); 40) D. geniculatus (male, Tibet); 41) D. sinicus (holotype); 42) D. elegans (male, New Guinea); 43) D. unidentatus (male, Java). Scale line 5 1.0 mm.