Embates todillofasciatus Prena, 2005

Prena, Jens, 2005, The Middle American species of Embates Chevrolat (Coleoptera: Curculionidae: Baridinae), Zootaxa 1100 (1), pp. 1-151 : 1-151

30. Embates todillofasciatus Prena sp. n.

(Fig. 114–116, 257)

Holotype male (dissected), Costa Rica , labeled: “ COSTA RICA, Prov. Alajuela, A.C.A./ San Ramón, Reserva Biol Alberto/ Brenes, Rio San Lorencito. 850m. / 19–26 MAR 1999. R. A. Zuñiga. / Manual (red libre) #52478”, INB0003025806 ( INBC).

Paratypes 12 (7 males, 5 females), labeled: “ COSTA RICA. Prov. San José,/ Cascajal, P.N.B.C., E. Zurquí,/ 0.500Km antes Túnel Zurquí, 1500m./ OCT 1990 – ABR 1991. G. Maass. / Intersección. L_N_535200_226800”, INB0003162434 ( INBC) ; “ COSTA RICA / F NEVERMANN”, “TURRIALBA/ 800 M/ SLG. SCHILD” ( NMNH) ; “ COSTA RICA, Prov. Cartago:/ P.N. Tapantí, Quebr. Segunda,/ 1200m, 9° 46' N 83° 47' W ”, “3.­ 5.5.2004 / leg. J. Prena ” ( JPPC) ; “ COSTA RICA, Puntarenas Prov. / Monteverde / 26 March 1987 / W.E. Steiner” ( NMNH) ; “ R. San Lorenzo, 1050m,/ R.F. Cord. Guanacaste / (Tenorio), Prov. Guan. / COSTA RICA. C./ Alvarado , Jun 1991,/ L­N­287800, 427600”, CRI000 602011 ( INBC) ; “ Costa Rica, HEREDIA: 5/ km N San Isidro, Cerro / Zurquí , 1800 m, 26.2.2000,/ leg. Prena ” ( JPPC) , same label except 1.4.2000 ( JPPC) ; “ COSTA RICA: Prov .

Heredia :/ 16km SSE La Virgen, 1050–/ 1150m, 10°16'N 84°05'W / 10.–14.III.2001 / INBio­OET­ALAS transect”; “handcollecting/ leg. J. Prena ”, INB0003209858 ( JPPC) , same label except 10.–14.IV.2001, INB0003209941 ( INBC) ; “ PANAMA, Chiriqui:/ Palo Alto 1400–1600m / 10. August 1995 / leg.: F. Oedegaard ” ( FOEC) ; “ PANAMA, Chir., Res. / For. La Fortuna , Quebra­/ da Aleman, 7­21­1995 / C.W.& L.B. O’Brien” ( CWOB) ; “ PANAMA, Chiriquí: Res. / La Fortuna, 1100 m / 8° 44'N 82°14'W / 21.– 25.3.2001, lg. Prena” ( HPSC) .

Description. Habitus: similar to Fig. 109, total length 4.2–5.8 mm (m=5.0, n=13). Color: integument piceous, basic vestiture of brown and some light­colored scales, yellow scales condensed in narrow subapical elytral fascia, small spots at interstria 6 (Fig. 114) and dorsolateral pronotal vitta, scales light yellow ventrally. Head: frontal fovea absent, rostrum slender, subcylindrical, curved (similar to Fig. 110), costate dorsomedially, basolateral margin slightly edged, length of rostrum ♂♂ 1.23–1.37 x (m=1.29, n=8), ♀♀ 1.26– 1.34 x (m=1.31, n=5) pronotal length, length of ante­antennal portion ♂♂ 0.38–0.40 x (m=0.39, n=8), ♀♀ 0.41–0.43 x (m=0.42, n=5) total rostral length, dorsal margin of antennal scrobe reaching rostral base before eye; funicular segment 2 much longer than 1, club oblong ovate. Pronotum: length 0.89–0.99 x (m=0.95, n=13) maximum width, subconical, widest in basal third, anterior portion tubulate; disk densely and shallowly punctate. Elytra: length 1.74–1.85 x (m=1.79, n=13) width at humeri, width 1.35–1.50 x (m=1.41, n=13) maximum pronotal width, sides subparallel in basal half, then increasingly narrowed toward apex, apices rounded conjointly, preapical callus costate, striae fine, punctures moderate, interstriae flat, 9 costate. Legs: tibiae nearly straight and parallel­sided, distally with cluster of cupreous hairs, tarsal claws arcuate and separate at base. Male: apex of aedeagus broadly rounded, middle sclerotized, anterolateral portion membranous (Fig. 115), body of aedeagus short, basal third angular in lateral view, apodemes approximately 2.8 x longer than body of aedeagus, flagellum thin, shorter than apodemes, transition to curved base gradual, basal appendage moderate, fused laterally with base of flagellum, projecting beyond base (Fig. 116).

Plant association. P. glabrescens (Prena 2), P. aequale (Prena 1), P. tenuimucronatum (Prena 1), P. dotanum (Prena 1).

Distribution. Costa Rica and Panama, evergreen montane forests of Atlantic side between 800 and 1800 m (Fig. 257).

Specific epithet. The name is a compound Latin participle derived from todillus (thin) and fasciatus (striped).

Discussion. Embates todillofasciatus is an uncommon species of the premontane forests in the Cordilleras Central and Talamanca. It belongs to the complex of slender species near E. pictipennis , and can be recognized easily by its color­pattern (Fig. 114). The narrow transverse line of light­colored scales derives from the post­macular element of a former elytral macula. The hypothesized ancestral color­pattern can be seen in the South American E. delicatulus (Hustache) ( Fig. 27 ).

Figures

FIGURES 26–33. Examples for dorsal color­patterns in species of Embates (for further explanations see in the text). 26–28, species with dark elytral macula preserved: 26, E. sagax, Peru, with elytral macula with circumferential line entire; 27, E. delicatulus, Bolivia, with elytral maculae and post­macular elements merged across suture, ante­macular element obliterated; 28, E. species #91, Colombia, with narrow ante­macular and wide post­macular elements. 29–33, species with dark elytral macula merged with basic vestiture: 29, E. obliquatus, Peru, with ante­macular element obliterated, post­macular element well­developed and humeral streak present; 30, E. rhombifer, Costa Rica, with ante­ and post­macular elements wide, fused with humeral streak and demarking rhomboid pseudo­macula on disk; 31, E. quadrilineatus, Guyana, with perfectly continuous vitta composed of ante­ and post­macular, humeral and apical elements; 32, E. species #75, Ecuador, with post­macular element obliterated, ante­macular and apical elements modified to fasciae; 33, E. species #101, Peru, with ante­macular, humeral and apical elements reduced to small spots, postmacular element disintegrated in two small spots.