Everettia planispira, Liew & Schilthuizen & Vermeulen, 2009

Liew, Thor-Seng, Schilthuizen, Menno & Vermeulen, Jaap Jan, 2009, Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo, Zoological Journal of the Linnean Society 157 (3), pp. 515-550 : 541

EVERETTIA PLANISPIRA SP. NOV.

Types: MALAYSIA: State of Sabah: Beluran district. Ulu Tungud Forest Reserve. East of Solonsong River , 400 m alt. (5°49 ′ N, 117°8 ′ E). Collected by M. Schilthuizen, T.- S. Liew, and A. van Til. Date: 14.viii.2004, holotype, BOR/MOL 3213, Figure 12E ; two paratypes, one in BMNH 20080633 and one in SP 13065 .

Etymology: Planispira . Latin, means flat whorls.

Material examined: SABAH – Danum Valley: BOR/ MOL 1041, vii.2001; BOR/MOL 1132, v.2000; BOR/ MOL 1139, v.2000; BOR/MOL 1142, iii.2001; BOR/MOL 1143, ix.2000. Tabin Wildlife Reserve: BOR/MOL 1146, v.2001. Kinabatangan: BOR/MOL 1308, Batu Keruak, iv.2002; BOR/MOL 1309, Batu Keruak, iv.2002; BMNH 1892.70.20.61, Gomantong, 1892. Mount Kinabalu: BOR/MOL 5358, eastern slope, 1952 m alt., iv.2004; BOR/MOL 5358, eastern slope, 2112 m alt., iv.2004.

Diagnostic characteristics: This species is easy to recognize by its small, shiny shell with flat apex and lack of sculpture on the upper and lower surface of the shell.

Description: Shell ( Fig. 12E ): small, rather thin, brownish, spire flat to slightly elevated, outer whorls flat to slightly rounded. Periphery rounded. Above the periphery, shell shiny with faded growth lines. Below the periphery, shell is smooth and shiny. Height up to 4.7 mm; width up to 8.3 mm; diameter of the first three whorls 0.8, 0.7, and 0.8 mm, respectively; number of whorls up to four and one-quarter; height aperture up to 3.5 mm; width aperture up to 4.8 mm. Genitalia: unknown. Animal: unknown.

Distribution and habitat: Primary forest 50–2000 m, but becoming very rare above 500 m. Sabah: Kinabatangan, Tabin, Danum valley, and Tawau (east coast of Sabah) ( Figs 5D , 9B ).

Remarks: This species has a shiny shell and is mainly found on the east coast of Sabah.

Figures

Figure 5. Distribution maps (Sabah). A, Everettia subconsul in the east and west coast lowland, Everettia jucunda in the west coast, and Everettia occidentalis sp. nov. in the western mountain range; B, Everettia klemmantanica in the central mountain ranges, Everettia interior sp. nov. in the interior area, Everettia paulbasintali sp. nov. in the east coast lowland, Everettia themis in the central mountain ranges; C, Everettia lapidini sp. nov. in the central mountain ranges, Everettia jucundior sp. nov. in the east coast lowland; D, Everettia monticola sp. nov. in the central mountain ranges, Everettia planispira sp. nov. in the eastern and northern parts of Sabah.

Figure 9. Distribution maps (Mount Kinabalu and Mount Tambuyukon). A, Everettia monticola sp. nov. and Everettia safriei sp. nov.; B, Everettia planispira sp. nov., Everettia jasilini sp. nov., and Everettia corrugata corrugata; C, Everettia dominiki sp. nov.; D, Everettia themis, Everettia subconsul, Everettia corrugata williamsi ssp. nov. and Everettia layanglayang sp. nov.

Figure 12. Shells. A, Everettia jasilini sp. nov.; B, Everettia monticola sp. nov.; C, Everettia dominiki sp. nov.; D, Everettia safriei sp. nov.; E, Everettia planispira sp. nov. Scale bar = 10 mm.