Ramírez-Ponce, Andrés & Morón, Miguel Angel, 2012, Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)

Ramírez-Ponce, Andrés & Morón, Miguel Angel, 2012, Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini), Zootaxa 3394, pp. 1-24

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Pachystethus     2-4
Pachystethus key    4-5
Pachystethus viduus     5-7
Pachystethus crassesculptus   new combination  7-9
Pachystethus nutans   new combination  11-13
Pachystethus nectoctenus   new species  13-15
Pachystethus matzapatlecus   new species  15-17
Pachystethus ixtacomitanus   new species  18-21