Ratcliffe, Brett C., 2002, A checklist of the Scarabaeoidea (Coleoptera) of Panama

Ratcliffe, Brett C., 2002, A checklist of the Scarabaeoidea (Coleoptera) of Panama, Zootaxa 32, pp. 1-48

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Passalini     5-6
Aesalini     5
Lucanini     5
Proculini     6
Ochodaeidae     7
Hybosoridae     7
Ceratocanthinae     7-8
Troginae     8-9
Athyreini     9
Bolboceratini     9
Aphodiini     9-10
Eupariini     10-11
Eurysternini     11
Rhypariini     11
Canthonini     12-13
Ateuchini     13-15
Coprini     15-16
Phanaeini     16-17
Onthophagini     17
Orphninae     18
Sericini     18
Melolonthini     18-19
Macrodactylini     19-20
Rutelini     20-24
Hopliini     20
Anomalini     24-26
Geniatini     26-27
Anoplognathini     26
Spodochlamyini     26
Cyclocephalini     27-31
Pentodontini     31-33
Oryctini     33-35
Phileurini     35-36
Dynastini     36-37
Agaocephalini     36
Gymnetini     38-39
Trichiini     39-40
Cremastocheilini     39
Cetoniini     39