A synopsis of the Euphaeidae (Odonata: Zygoptera) of Vietnam, with descriptions of two new species of Euphaea Author Phan, Quoc Toan Author Kompier, Tom Author Karube, Haruki Author Hayashi, Fumio text Zootaxa 2018 2018-01-24 4375 2 151 190 journal article 30935 10.11646/zootaxa.4375.2.1 c8d905a3-01d4-4d5f-a904-fad72add2ae1 1175-5326 1158445 7A4595B8-9FDF-41A9-9C39-A8A4A83FF302 Euphaea ochracea Selys, 1859 ( Fig. 17a–e ) Euphaea ochracea : Selys (1859) , p. 443; Selys (1891) , pp. 849–490; Laidlaw (1902) , p. 87; Laidlaw (1903) , p. 193; Martin (1904) , p. 218 [‘ Tonkin , Annam’; first record from Vietnam ]; Lieftinck (1954) , p. 17; Schmidt (1964) , p. 153; Asahina (1970) , p. 112; Hämäläinen et al. (1996) , p. 5; Kitagawa (1997) , p. 13; Yokoi & Souphanthong (2005) , p. 38 (species list); Do & Dang (2007) , p. 36, distribution map [Records from Lai Chau , Ha Tinh and Nghe An Provinces]; Zhang (2010) , p. 25 (species list); Dow et al. (2016) , p. 7; Hämäläinen (2017) , p. 11; Khan (2017a) , p. 15 (species list). Euphaea ochlacea (sic): Asahina (1996) , p. 190 [Records from Lai Chau Province ]. Allophaea brunnea : Fraser (1933) , pp. 110, 123. Allophaea ochracea : Fraser (1934) , pp. 96–98, Fig. 3a (wings); Fraser (1942) , p. 98. Allophaea ochracea ochracea : van Tol & Rozendaal (1995) , pp. 99–100. [Records from Lai Chau, Ha Tinh and Nghe An Provinces]. Euphaea ochracea ochracea : Laidlaw (1931) , p. 180; Asahina (1961) , p. 210; Asahina (1964) , p. 114; Asahina (1977) , p. 173; Yokoi & Mitamura 1995 , p. 7 (species list); Yokoi (1999) , p. 3 (species list); Mitra (2002) , Figs 27, 39–40; Yokoi & Souphanthong (2004) , p. 54 (species list); Sasamoto et al . (2011) , p. 2 (species list). Euphaea ochracea brunnea: Lahiri (1987) , p. 40, Figs 75–76, 111–112. Pseudophaea ochracea : Williamson (1904) , pp. 181–182; Laidlaw (1924) , p. 297. Materials examined. [ Vietnam ] 1♂ , Nam Bung , Yen Bai Prov., 1.VI.2014 , T. Kompier leg. (TKC) ; 1♂ , Nam Bung , Yen Bai Prov., 10.V.2014 , T. Kompier leg. (TKC) ; 3♂ , Vu Quang National Park , Ha Tinh Prov., 7.IX.2015 , Q.T. Phan leg. ( PQTC ) ; 1♂ , A Luoi Nature Reserve , Thua Thien-Hue Prov., 18.IX.2015 , Q.T. Phan leg. ( PQTC ) ; 3♂ , Bhalee , Tay Giang District , Quang Nam Prov., 18.IX.2015 , Q.T. Phan leg. ( PQTC ) ; 2♂ , Tr’hy , Tay Giang District , Quang Nam Prov., 20.VII.2017 , Q.T. Phan leg. ( PQTC ) ; 1♂ , Quang Nam Prov., 20.VI.2016 , T. Kompier leg. (TKC) ; 1♂ 1♀ , Mang Den , Kon Plong District , Kon Tum Prov., 20.IX.2015 , F. Hayashi leg. (FHC). [ China ] 2♂ 2♀ , Xishuangbanna , 1300 m , Yunnan Prov. , 3.IX.2006 , X. Yu leg. ( PQTC ). [ Laos ] 1♂ 1♀ , Xam Neua , Houaphan Prov. , 9.VI.2017 , X. Liu leg. (FHC). [ Malaysia ] 2♂ 5♀ , Ipoh , Perak state , Malaysia , 28.IV.2012 , F. Hayashi leg. (FHC) ; 1♂ , Penang Hill , Penang , Malaysia , 7.III.2014 , Y. Kamimura leg. (FHC). Other materials confirmed by field observations. [ Vietnam ] Some individuals, Yen Bai Prov., IV , V , VII.2014 , IV , V , VI.2015 , VII.2016 , T. Kompier ; 20 exs , Hoa Binh Prov., 12.V.2015 , T. Kompier ; Some individuals , Thanh Hoa Prov., V , VI.2015 , T. Kompier ; Some individuals , Quang Nam Prov., IX.2015 , VI.2016 , T. Kompier ; 2 exs, 4.VI.2017 , Quang Binh Prov., 15 exx, 3.IX.2016 , T. Kompier ; Quang Binh Prov., T. Kompier; 25 exs, 8.IX.2016 , Quang Binh Prov., T. Kompier. Notes. This species is distributed in northern and central Vietnam . Male wings are pale orange colored, but the apices are hyaline. Bodies of male and female are shown in Figure 17a and 17b , respectively. Asahina (1977) noted that male specimens taken in Malaysia and southern Thailand have an entirely pale brownish [sic] HW, but that in Laotian males, the brownish [sic] area of HW is restricted before the pterostigma, although a clear geographical differentiation using this character seems difficult. We also confirmed this difference between Malaysia and Laos . In Vietnam and China , all male specimens examined have a HW similar to that observed in Laotian males ( Fig. 17a ). Male genital ligula has laterally curved flagella typically seen in other Euphaea species, but these are relatively narrow and elongated ( Fig. 17c ). Distribution. Vietnam ( Lai Chau , Yen Bai , Hoa Binh , Thanh Hoa , Nghe An , Ha Tinh , Thua Thien-Hue , Quang Nam and Kon Tum Provinces), China ( Yunnan Province ) ( Zhang 2010 ), Laos ( Phongsali , Luang Namtha , Oudomxay , Houaphan , Vientiane and Attapeu Provinces) ( Fraser 1933 , Asahina 1977 , Yokoi & Mitamura 1995 , Yokoi 1999 , Yokoi & Souphanthong 2004 , 2005 , Sasamoto et al . 2009, this study), Thailand (most of the country, except Central Plains and North-east) ( Hämäläinen 2017 ), Myanmar ( Shan and Bago Provinces [ Schmidt 1964 ], unknown in detail [ Asahina 1970 ]), Peninsular Malaysia ( Johor , Kelantan , Pahang , Penang , Perak , Perlis and Selangor States) ( Lieftinck 1954 , Laidlaw 1902 , 1903 , 1924 , 1931 , Fraser 1942 , Hämäläinen et al. 1996 , Kitagawa 1997 , Dow et al. 2016 ), Nepal (Kathmandu Valley) ( Vick 1989 ), Bangladesh ( Chittagong and Sulhet divisions) ( Khan 2017a ), India ( Meghalaya [ Lahiri 1987 ], Arunachal Pradesh , Assam , Manipur , West Bengal and Mizoram [ Mitra 2002 ]).