Taxonomic review of tropical western Atlantic shallow water Drilliidae (Mollusca: Gastropoda: Conoidea) including descriptions of 100 new species
Author
Fallon, Phillip J.
text
Zootaxa
2016
4090
1
1
363
journal article
51606
10.11646/zootaxa.4090.1.1
e6b35f5a-435e-4473-b29e-1e4d842f84b0
1175-5326
263299
203BAC25-B542-48FE-B5AD-EBA8C0285833
Fenimorea paria
(Reeve, 1846)
,
nomen dubium
(
Plate 91
)
Pleurotoma paria
Reeve, 1846
a: pl. 36, fig. 334: Reeve (1846c: 5);
Drillia
(
Crassispira
)
paria
(Reeve, 1846)
: H. & A. Adams (1853: 91 [list]).
Not this species, or unconfirmed reports
.
Drillia
(
Clavus
)
fucata
(Reeve, 1845)
, as a synonym of: Tryon (1884: pl. 11, fig. 93).
Pleurotoma fucata
Reeve, 1845
, as a synonym of: Paetel (1888).
Drillia fucata
var.
paria
(Reeve, 1845)
: Dall (1889a: 89); Dall (1889b: 98, 99); Dall (1903: 98).
Drillia paria
(Reeve, 1846)
: Hidalgo (1904: 116); Rios (2009: 307).
Cymatosyrinx furcata
[sic]
paria
(Reeve, 1846)
: Johnson, C. W. (1934: 136).
Fenimorea fucata paria
(Reeve, 1846)
: Fargo (1953: 369); Powell (1966: 90); Porter (1974: 252), cites Dall’s Cape Fear, North Carolina locality.
Splendrillia
(
Syntomodrillia
)
fucata
(Reeve, 1845)
, as a synonym of: Abbott (1974: 275).
Pleurotoma paria
Reeve, 1846
: Maes (1983: 313 [on confusion with
Drillia cydia
(Bartsch, 1943)
]).
Clathrodrillia paria
(Reeve, 1846)
: Rios (1983); Rios (1985:136; pl. 46, fig. 620); Rios (1994: 159; pl. 53, fig. 711); Williams (2005, 2006, 2009: species 1544).
Turris
(
Drillia
)
paria
(Reeve, 1845)
: Faustino (1928: 314 [Zamboanga,
Philippines
]).
Types
.
Two
syntypes
: 13.4 x 5.8 & 16.1 x
6.2 mm
(NHMUK 1963925).
Type
locality.
Unknown.
PLATE 91.
Fenimorea paria
(Reeve, 1846)
. Figs. 1–2: syntypes, locality unknown, (NHMUK 1963925). Fig. 1: ventral, lateral & dorsal views of smaller syntype; Fig 2: ventral view of larger syntype.
Description.
Shell
. The best preserved
syntype
is a mature shell, 13.4 x
5.8 mm
, fusiform, truncated anteriorly; with a large body whorl, approximately 61% of the shell’s total length. The shell has a drill hole and is missing its protoconch; 9–10 whorls remain.
Protoconch
missing on the
syntypes
.
Axial sculpture
of low, broad ribs that extend from suture-to-suture on the spire whorls, but fade on the body whorl at the anterior fasciole. Ribs number 14 on the penultimate and 10 to the varix on the body whorl, are reduced and arcuate in the sulcus.
Spiral sculpture
with dense microscopic spiral grooves cut by growth striae, forming a pattern typical of
Fenimorea
.
Sulcus
is demarcated by the reduction in the size of the ribs and the presence of spiral grooves.
Varix
is large, broad, hump-like; ⅓-turn from the edge of the outer lip. The excavation around the large drill hole just above the varix gives the varix the appearance of being more protuberant than it is (Pl. 91,
Fig. 1
, lateral view).
Outer lip
thin, chipped, with 4 axial folds; no stromboid notch present, or perhaps lost in the wear of the outer lip.
Anal sinus
on whorl shoulder, a deep U-shaped notch bordered by a lobe of the inner lip on the parietal side, and by a projection of the outer lip. Edge of sinus flared out such that it appears spout-like.
Inner lip
erect anteriorly along the canal, recumbent elsewhere, widest on parietal wall, and developed into a lobe posteriorly, at the roof of the aperture.
Anterior canal
short, open, directed to the left when viewed ventrally, very slightly notched; anterior fasciole not swollen.
Color
white with a few faint brownish-yellow maculation—appearing band-like on the last whorl, otherwise randomly placed on whorl shoulders.
Remarks.
Nomenclature.
Fenimorea paria (Reeve, 1846)
has a history of uncertain placement. It has been allied with
F. fucata
(Reeve, 1845)
variously as a variety, subspecies or a junior synonym, or white form of
F. fucata
. The
types
exhibit the characteristics of
Fenimorea
; the smaller specimen even possesses the brown band and random brown patches of
F. fucata
, albeit very faint. They are however in too poor a condition to make a definite identification. If
F. paria
is conspecific with
F. fucata
, the latter name would have priority. All-white forms of
F. f u ca t a
-like specimens
F. paria
as
F. p ar i a
is not all white, and all-white forms of several different species of
Fenimorea
are known to exist. Neither are there other unique diagnostic characteristics that would support
F. pa r i a
as distinct from other named species. No populations of
F. fucata
-like species have been found that are nearly all white with faint brown maculation as exhibited by
F. p ar i a
. Inasmuch as there is doubt at this time that it is a valid taxon and not just worn specimens of an already described
F. f uc a t a
-like species and because no similar specimens have been observed, it is best treated as a
nomen dubium
and use of the name avoided until further evidence of its identity emerges.