The fast-running flies (Diptera, Hybotidae, Tachydromiinae) of Singapore and adjacent regions Author Grootaert, Patrick Author Shamshev, Igor V. text European Journal of Taxonomy 2012 2012-01-06 5 1 162 journal article 10.5852/ejt.2012.5 e809cd3a-36bb-413d-9f19-70837d7db34b 3715167 B69F002A-C1A0-439D-9477-62BFA87DEAD7 Drapetis pantai sp. nov. Figs 92 , 105-107 Diagnosis Rather small species; antenna brownish yellow; palpus brownish; legs brownish yellow, hind tarsomere 1 with some longer ventral spinules near base; halter pale. Description Male LENGTH. Wing 1.5-1.6 mm . HEAD. Black. Occiput subshining, finely pollinose. Two long vertical bristles present. Frons subshining, broadened towards ocellar tubercle. Anterior ocellars long, yellow; posterior ocellars minute. Antenna brownish yellow, postpedicel somewhat darker; pedicel with moderately long lower seta; postpedicel with lower margin somewhat convex, rather long, 2.5-2.8 times longer than wide; stylus nearly 3.0 longer than postpedicel and about 1.5 times longer than scape, pedicel and postpedicel combined. Palpus brownish, with moderately long subapical seta. THORAX. Black. Anepisternum pubescent. Prothoracic episternum with 1 long upturned bristle just above fore coxa and some setulae on upper part. Postpronotal bristle not prominent. Mesonotum with 2 notopleurals, 1 postsutural supra-alar, 1 postalar and 4 scutellars (inner pair very long, cruciate; outer pair very short); scutum entirely covered with short, pale uniform (except for 1 pair of long prescutellars) setulae. LEGS. Brownish yellow with hind femur, all tibiae and tarsi darker in different extent (in darker specimens hind femur and all tibiae brownish). Coxae and trochanters with unmodified setation. Fore femur somewhat thickened. Femora and tibiae mostly with inconspicuous setation. Mid femur with 1 anterior subapical bristle. Hind femur slender, with 3-4 longer anteroventral subapical setae ( Fig. 92 ). Hind tibia with slightly prominent apical projection. Hind tarsomere 1 with some longer ventral spinules near base. WING. Normally developed, hyaline, covered with uniform microtrichia; veins yellowish to brownish yellow. Costal vein with short setulae along anterior margin. Costal bristle short, brownish. Costal index: 50/16/47. Vein Rs originating before middle of cell r 1 (closer to humeral crossvein), nearly as long as crossvein bm-cu. R 1 thickened, meeting costa about middle of wing. R 2+3 evenly bowed toward costa, meeting costa beyond middle of wing. R 4+5 and M 1+2 convergent in apical part. R 4+5 slightly sinuate. A 1 absent. Crossvein bm-cu somewhat oblique. Crossvein r-m near middle of cell bm. Halter pale. Figs 105-107. Drapetis pantai sp. nov. , ♂ terminalia. 105 . right surstylus, dorsal view. 106 . epandrium and cerci, dorsal view. 107 . left surstylus, lateral view. ABDOMEN. With tergite 1 pale yellow, tergites 2-7 brownish, rectangular, of subequal width, with scattered setulae, tergites 3 and 4 with squamiform setae laterally; segment 8 brown. Sternite 2 divided, sternites 3 and 4 with excision posteriorly. Gland-like structures present between tergites 3-4 and 4-5. TERMINALIA. Small, black ( Figs 105-107 ). Cerci separated; left cercus broad ( Fig. 106 ), subrectangular, with numerous unmodified setae of different lengths; right cercus greatly reduced, represented as narrow, bacilliform sclerite bearing several short setulae. Epandrium completely divided. Left epandrial lamella fused to hypandrium, lacking setae in apical part. Left surstylus ( Fig. 107 ) bilobed; upper lobe rather digitiform, with numerous unmodified setae; lower lobe somewhat shorter and broader, truncate, with several unmodified setae. Right epandrial lamella ( Fig. 106 ) subtriangular, covered with numerous unmodified setae of different lengths. Right surstylus bilobed ( Fig. 105 ); upper lobe broad, with several long unmodified setae; lower lobe somewhat narrower and longer, with unmodified setae. Hypandrium reduced, lacking setation. Phallus short. Two rod-shaped apodemes present. Female Unknown. Material examined Holotype SINGAPORE : Chek Jawa, 15 December 2005 , mangrove, sweeping (reg. 25452, leg. P.G.; in ZRC). Paratypes SINGAPORE : 1 ♂ , Chek Jawa, 15 December 2005 , mangrove, sweeping (reg. 25452, leg. P.G.). MALAYSIA : 3 ♂♂ , Pulau Tioman, Juara, 20 July 2005 , river, sweeping (reg. 25244, leg. P.G.). Etymology The specific name 'pantai' means beach or coast in Malay. Distribution Malaysia , Singapore . Habitat and seasonal occurrence This species is found in mangroves, recorded in July and December. Remarks This species resembles representatives of the D. assimilis and D. convergens groups ( sensu Kovalev 1972 ). However, the new species is likely a member of a hitherto unrecognised group within Drapetis , defined on the basis of the following features: prothoracic episternum with 1 long upturned bristle just above fore coxa, tergites 3 and 4 with squamiform setae.