' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae) Author Jałoszyński, Paweł text Zootaxa 2014 3828 1 1 76 journal article 36382 10.11646/zootaxa.3828.1.1 8c1cda9b-0d82-41f3-a1c1-d8eb574a1ba7 1175-5326 286513 B2FEAE60-7E51-45FA-A38F-930A084A5AAA Horaeomorphus verus sp. n. ( Figs. 26 , 31–32 , 35–36 , 193 b) Type material studied . Holotype : AUSTRALIA : ♂: three labels: " 16.05S 145.28E QLD / CT2 10m , 2 km SSW / Cape Tribulation / 1-28 Feb 1996 / L.Umback / FI Trap JCU" [white, printed], " ANIC / Specimen" [green, printed], " HORAEOMORPHUS / verus m. /, det. P. Jałoszyński, 2013 / HOLOTYPUS " [red, printed] ( ANIC ). Diagnosis. Male: BL> 2 mm ; pronotum with three antebasal pits connected by distinct transverse groove. Male: aedeagus in ventral view with apical part broadly rounded. Description. Body ( Fig. 26 ) flattened, elongate and slender, with moderately long appendages, BL 2.05 mm ; cuticle glossy, body uniformly dark brown with slightly lighter appendages, vestiture light brown. Head ( Fig. 26 ) broadest at eyes, HL 0.33 mm , HW 0.41 mm ; tempora slightly longer than eyes, strongly curving posteromesally; vertex strongly transverse, more convex on sides than in middle, with distinctly concave posterior margin; frons subtrapezoidal, posteriorly confluent with vertex, anteriorly gradually lowering toward clypeus; supraantennal tubercles prominent, each sharply demarcated from median part of frons and indistinctly demarcated laterally from vertex, each accompanied by pair of deep pits near posteromesal margin. Eyes moderately large, bean-shaped, weakly protruding laterally from the head silhouette, finely faceted. Punctures on head dorsum fine and sparse, inconspicuous; setae short, sparse and suberect, except for sparse and short erect bristles on tempora. Antennae ( Fig. 26 ) slender, AnL 0.88 mm ; antennomeres I–VI each distinctly elongate, VII about as long as broad, VIII–X strongly transverse, XI shorter than IX–X together, about 1.5x as long as broad, with indistinctly pointed apex. Pronotum ( Fig. 26 ) distinctly elongate and weakly convex, broadest near anterior third, PL 0.58 mm , PW 0.53 mm ; anterior and lateral margins rounded; sides moderately strongly convergent posteriorly; hind pronotal corners distinct but obtuse and blunt; posterior margin weakly arcuate; base of pronotum with three small but deep and distinct pits connected by sharply marked transverse groove. Punctures on pronotal disc small but distinct, in middle of disc separated by spaces about as wide as 1-1.5x diameters of punctures; setae moderately dense, thin, short and suberect, sides of pronotum with bristles. Elytra ( Fig. 26 ) oval, as convex as pronotum, broadest slightly behind anterior third, EL 1.15 mm , EW 0.75 mm , EI 1.53; basal impressions short; humeral calli developed as elongate protuberances distinctly demarcated mesally from basal part of elytra; elytra with indistinct circumsutural impression near anterior third, small area behind impression slightly raised; elytral apices rounded together. Punctures on elytra unevenly distributed, those on circumsutural impression larger and deeper than those on pronotum, sharply marked and separated by spaces as wide as 0.5–1 diameters of punctures, punctures reducing in diameter, depth and sparser toward elytral margins; setae moderately long and dense, suberect. Hind wings well developed, about twice as long as elytra. Legs ( Fig. 26 ) moderately long and slender, without modifications. Aedeagus ( Figs. 31–32 , 35–36 ) moderately stout, AeL 0.40 mm , in ventral view apical part subtrapezoidal, apical margin rounded in middle; internal armature darkly sclerotized and complicated, symmetrical, with long subapical median tubular structure and bell-shaped central complex of sclerites; parameres slender, not exceeding apex of median lobe, with long apical setae. Female . Unknown. Etymology. The Latin adjective verus (real, true, genuine) was chosen as a specific epithet, to underline that this is the first and only true Australian Horaeomorphus originally placed in a correct genus. Distribution ( Fig. 193 b). North-eastern Australia (northern Queensland). Remarks. Horaeomorphus verus has the aedeagus similar to that of H. anthicoides , but clearly differs in the external structures; e.g., the body length, proportions of body parts, the sculpture of pronotal base and much longer and denser setae on the dorsum. For characters separating this species from Oriental congeners see the Remarks section at the redescription of H. anthicoides .