Bryozoa of Floridan Oculina reefs Author Judith L Winston text Zootaxa 2016 4071 1 1 81 journal article 51549 10.11646/zootaxa.4071.1.1 8ffe5a88-06f7-44af-95aa-5fd49e0302c4 1175-5326 260490 D47C792F-E91D-40A6-ABB7-FA7810578562 Plesiocleidochasma cleidostomum (Smitt, 1873) comb. nov. ( Fig. 35 ; Table 34 ) Lepralia cleidostoma Smitt, 1873: 62 , pl. 11, figs 217–219. Lepralia porcellana : Osburn 1914: 209 . Hippoporina cleidostoma : Canu & Bassler 1928a: 104 , pl. 9, fig. 7, pl. 32, fig. 6, text-fig. 18. Hippoporina porcellana : Osburn 1940: 428 ; Shier 1964: 633. Cleidochasma porcellanum : Cheetham & Sandberg 1964: 1032 , text-fig. 35; Long & Rucker 1970: 19, fig. 4.2; Winston 1982: 147, fig. 80. Schedocleidochasma porcellanum : Winston 2005: 87 , figs 244–246. TABLE 34. Measurements in mm of Plesiocleidochasma cleidostomum (Smitt, 1873) .
Lz Wz Lo Wo Lov Wov Lav Wav
N 18 18 18 18 12 12 1 1
Mean 0.428 0.321 0.124 0.095 0.182 0.240 0.126 0.054
SD 0.043 0.031 0.009 0.011 0.021 0.019 ------ ------
Min 0.360 0.288 0.108 0.081 0.144 0.148 0.126 0.054
Max 0.522 0.396 0.144 0.126 0.216 0.270 0.126 0.054
Material examined. MCZ # 90, syntype of Lepralia cleidostoma , May 1, 1868 , Cast No. 7, 26 fms [off Sombrero Key], L. Pourtales; VMNH no. 70652, 70653, 70654; USNM no. 1283259. Description. Colonies encrusting, in unilaminar patches to multilaminar mounds. Primary layer of zooids regularly hexagonal; frontally budded zooids irregularly polygonal. Frontal shield smooth to sparsely granular, flat over most of frontal surface ( Fig. 35 B), raised around orifice, imperforate except for a few marginal pores. Orifice cleithridiate, rounded distally with proximally directed condyles, and narrow U-shaped sinus. Single or paired adventitious avicularia may occur on umbo proximolateral to orifice; rostra and mandibles acutely triangular ( Fig. 35 C). Ovicell hyperstomial, prominent and delicately striated when newly formed, becoming granular and embedded in secondary calcification with age; imperforate, with frontal area in form of a thin curved-edged plate (labellum) between two lateral slits, leaving narrowly transverse opening ( Fig. 35 A, B). Embryos reddish-pink. Discussion . Floridan Plesiocleidochasma cleidostomum is part of the ‘ Cleidochasma porcellanum ’ species complex, widely distributed in tropical and subtropical habitats, including the Western Atlantic (see Winston 2005). Soule, Soule & Chaney (1991) and Cook & Bock (1996) revised the genera and species included in the Cleidochasmatidae . Winston (2005) placed Smitt’s species in Schedocleidochasma , but Berning (2012) recently synonymized Schedocleidochasma under Plesiocleidochasma , a decision followed here. Distribution . Cape Hatteras to Florida, Caribbean, and Gulf of Mexico .