Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina) Author Taphylininae Author Iochini Author Aorothiini Author Thiini Author Latyprosopini Author Taphylinini Author Mblyopinina Author Nisolinina Author Yptiomina Author Hilonthina Author Herman, Lee H. BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY Number 265, pages 2441 - 3020 Issued July 18, 2001 (Parts I-VII) text Bulletin of the American Museum of Natural History 2001 2001-07-18 2441 265 2435 3021 http://dx.doi.org/10.1206/0003-0090.265.1.5 journal article 10.1206/0003-0090.265.1.5 0003-0090 10112575 tarsalis Sharp, 1876 , see: nanus Erichson, 1839 . testaceus Bernhauer, 1932b: 145 ( Diochus ; Type locality: Mulongo: Malinge; Niunzu). — Scheerpeltz, 1933: 1324 ( Diochus ; catalog). - DISTRIBUTION: Congo . testaceus Coiffait, 1981 , see: montguilloni Coiffait, 1987 . thoracicus Casey, 1906 , see: schaumii Kraatz, 1860 . ventralis Coiffait, 1968: 160 ( Diochus ; Type locality: Gabon : Belinga ). - DISTRIBUTION: Gabon . vicinus Sharp, 1876 , see: nanus Erichson, 1839 . vilis Sharp, 1885 , see: nanus Erichson, 1839 . Tribe Maorothiini Maorothiini Assing, 2000: 16 (genera included: Maorothius ; characters; phylogenetic analysis of tribal relationships). TYPE GENUS: Maorothius Assing , fixed by original designation. Maorothius [20 species; Australian Region ] Maorothius Assing, 2000: 22 , 58 (species included: adustus , hamifer , setiger , longispinosus , brevispinosus , insulanus , pubescens , brookesi , effeminatus , hammondi , pectinatus , puncticeps , coalitus , tridens , volans , dispar , brouni , torquatus , tonsor , solus ; phylogenetic analysis; key to species). TYPE SPECIES: Othius adustus Broun , fixed by original designation. adustus Broun, 1880: 101 ( Othius ; Type locality: Parua; [ Note: See lectotype designation by Assing, 2000]). — Bernhauer and Schubert, 1914: 317 ( Othius ; catalog). Assing, 2000: 24 ( Maorothius ; lectotype designation: Parua , Whangarei , New Zealand ; characters; notes; New Zealand ) . - DISTRIBUTION: New Zealand . angustus Broun, 1893 , see: brouni Steel, 1949 . brevispinosus Assing, 2000: 32 , 63 ( Maorothius ; Type locality: AK: Ahuroa forest reserve). - DISTRIBUTION: New Zealand . brookesi Cameron, 1952c: 42 ( Othius ; Type locality: New Zealand : North Island : Maku River , Bay of Plenty ). — Assing, 2000: 36, 63 ( Maorothius ; characters; notes; New Zealand ). - DISTRIBUTION: New Zealand . brouni Steel, 1949a: 270 ( Gyrohypnus ; replacement name for angustus Broun ). — Assing, 2000: 51, 63 ( Maorothius ; characters; notes; New Zealand ). - DISTRIBUTION: New Zealand . angustus Broun, 1893: 1026 ( Othius ; [ preoccupied ]; Type locality: Taieri). — Bernhauer and Schubert, 1914: 317 ( Othius ; catalog). — Steel, 1949a: 270 ( Gyrohypnus ; synonym of brouni ). — Assing, 2000: 51 ( Maorothius ; synonym of brouni ). coalitus Assing, 2000: 45 ( Maorothius ; Type locality: New Zealand : NN, Beebys Knob , 1220 m ). - DISTRIBUTION: New Zealand . dispar Assing, 2000: 49 ( Maorothius ; Type locality: L. Rotoroa, 1200'). - DISTRIBUTION: New Zealand . effeminatus Assing, 2000: 39 , 63 ( Maorothius ; Type locality: N. Z.: Whangarei). - DISTRIBUTION: New Zealand . hammondi Assing, 2000: 41 ( Maorothius ; Type locality: New Zealand : WO: Kauri Flat, 14 km . SW Waingaro ). - DISTRIBUTION: New Zealand . insulanus Assing, 2000: 33 ( Maorothius ; Type locality: New Zealand : Lt Barrier ). - DISTRIBUTION: New Zealand . hamifer Assing, 2000: 27 ( Maorothius ; Type locality: Waipu). - DISTRIBUTION: New Zealand . longispinosus Assing, 2000: 30 , 63 ( Maorothius ; Type locality: N-Zealand: Aukland [sic]). - DISTRIBUTION: New Zealand . pectinatus Assing, 2000: 42 ( Maorothius ; Type locality: New Zealand : ND, Russell SF , Runaruku). - DISTRIBUTION: New Zealand . pubescens Assing, 2000: 34 ( Maorothius ; Type locality: Waitotara, Wanganui). - DISTRIBUTION: New Zealand . puncticeps Broun, 1894: 419 ( Othius ; Type locality: Mount Pirongia ). — Bernhauer and Schubert, 1914: 318 ( Othius ; catalog). — Assing, 2000: 43 ( Maorothius ; characters; notes; New Zealand ). - DISTRIBUTION: New Zealand . setiger Assing, 2000: 29 , 62 ( Maorothius ; Type locality: N. Zeal.: Waipoua For., Waipoua Strm., 70 m ). - DISTRIBUTION: New Zealand . solus Assing, 2000: 57 ( Maorothius ; Type locality: New Zealand WN, Kowhai Park , Brooklyn ). - DISTRIBUTION: New Zealand . tonsor Assing, 2000: 56 , 63 ( Maorothius ; Type locality: New Zeal.: S Isl., Banks Penins). - DISTRIBUTION: New Zealand . torquatus Assing, 2000: 55 ( Maorothius ; Type locality: Oaro, N. Canterbury ). - DISTRIBUTION: New Zealand . tridens Assing, 2000: 46 ( Maorothius ; Type locality: WN: Desert Rd, S . Turangi, Mangatawai Stm. ). - DISTRIBUTION: New Zealand . volans Assing, 2000: 47 ( Maorothius ; Type locality: Maungatapere Bush). - DISTRIBUTION: New Zealand . Tribe Othiini Othiini Thomson, 1859 . — Reitter, 1908: 102 (characters). — Portevin, 1929: 367 (characters; key to French genera). — Scheerpeltz, 1933: 1297 (synonym of Xantholinini ). — Jeannel and Jarrige, 1949: 371 (cave species). — Jarrige, 1952a: 131 (characters). — Coiffait, 1956a: 37 (review of French taxa; characters). — Lohse, 1964: 166 (characters; key to central European species). — Scheerpeltz, 1968a: 63 (catalog of Austrian species). — Coiffait, 1972: 127, 326 (tribe of Xantholininae ; characters; key to genera and species of western Palaearctic region). — Muona, 1979: 16 (list of Scandinavian species). — Smetana, 1982: 35 (characters; revision of and key to genera and species of North America north of Mexico ). — Bordoni, 1982: 378 (key to genera of Italy ). — Newton, 1985: 208 (notes on austral distribution and taxonomic notes). — Outerelo and Gamarra, 1985: 43 (key to genera of Iberian Peninsula). — Newton and Thayer, 1992: 64 (tribe of Staphylininae ). — Klimaszewski, Newton, and Thayer, 1996: 150 (list of New Zealand genera; number of New Zealand species). — M. Hansen, 1996: 108 (list of Danish species). — Downie and Arnett, 1996: 396 (characters). — Newton, Thayer, Ashe, and Chandler, 2000: 328, 390 (key to Nearctic genera). Othiides Thomson, 1859: 26 (genera included: Othius , Gyrohypnus ). TYPE GENUS: Othius (see Newton and Thayer, 1992: 64). Othii Thomson, 1859. — Casey, 1906: 359, 422 (characters; key to genera). Othiina Thomson, 1859. — Jakobson, 1909: 498 (characters). Atrecini Hatch, 1957: 172, 231 (key to genera of Pacific Northwest; genera included: Atrecus , Parothius ). TYPE GENUS: Atrecus (see Newton and Thayer, 1992: 64). — Smetana, 1982: 35 (synonym of Othiini ). — Newton and Thayer, 1992: 64 (synonym of Othiini ). Atrecus [14 species; Nearctic and Palaearctic Regions ] Atrecus Jacquelin du Val, 1856: 31 (species included: pilicornis ). TYPE SPECIES: Staphylinus pilicornis Paykull , fixed by original designation by monotypy. — Schaum, 1859: 27 (synonym of Baptolinus ). — Gemminger and Harold, 1868: 606 (synonym of Baptolinus ). — Jacquelin du Val, 1868: 65 (catalog; European species). — Fauvel, 1873b: 86 [= 1873a: 372] (synonym of Baptolinus ). — Mulsant and Rey, 1877a: 237 (synonym of Baptolinus ). — Ganglbauer, 1895: 472 (synonym of Baptolinus ). — Bernhauer and Schubert, 1914: 316 (synonym of Baptolinus ). — Tottenham, 1949: 370, 417 ( type species: affinis Paykull = pilicornis Jacquelin du Val ; notes; checklist of British species). — Blackwelder, 1952: 69 ( type species: pilicornis ). — Hatch, 1957: 232 (characters; key to species of Pacific Northwest). — Smetana, 1958: 117 (characters; habitus of one species; type species: pilicornis ; key to Czechoslovakian species). — Moore, 1963a: 94 (characters). — Székessy, 1963: 51 (characters). — Lohse, 1964: 167 (synonym of Baptolinus ). — Pototskaia, 1967: 47 (larval characters). — Tikhomirova, 1973a: 180 (checklist of species of USSR). — Moore and Legner, 1975: 72 (catalog; North America north of Mexico ). — Shilov, 1975a: 375 [= 1975b: 97] ( Atrecus ; Komi region of Russia ). — Pototskaia, 1976: 171 (larval characters). — Pope, 1977: 28 (list of British species). — Moore and Legner, 1979: 55 (characters; notes). — Muona, 1979: 16 (list of Scandinavian species). — Smetana, 1982: 36 (characters; revision of and key to species of North America north of Mexico ). — Bordoni, 1982: 397 (synonym of Baptolinus ). — Shibata, 1983: 85 (checklist of species of Japan ). — Muona and Viramo, 1986: 14 (list of species of northeastern Finland ). — Lohse and Lucht, 1989: 159 (valid name). — Angelini, 1991: 190 (list of species from high elevations of Calabria , Italy ). — Campbell and Davies, 1991: 123 (checklist of species; Alaska and Canada ). — Silfverberg, 1992: 19 (list of species of Fennoscandia, Denmark , and the Baltic States). — Welch, 1993: 228 (cited as Atrechus ; ovariole structure and development). — Janák, 1993a: 47 (list of species of Czech Republic and Slovakia ). — Ciceroni and Zanetti, 1995: 26 (list of species of Italy ). — M. Hansen, 1996: 108 (list of Danish species). — M. Hansen, Mahler, Palm, and Pedersen, 1996: 243 (additions to list of Danish species). — Downie and Arnett, 1996: 396 (characters; key to species of Northeastern North America). — Assing, 1998c: 349 (key to species of Palaearctic region). — Newton, Thayer, Ashe, and Chandler, 2000: 390 (5 Nearctic species; notes; characters in key). Baptolinus Kraatz, 1857d: 659 (species included: alternans , pilicornis ). TYPE SPECIES: Staphylinus pilicornis Paykull , fixed by subsequent designation by Tottenham, 1945: 70. — Redtenbacher, 1857: 982 (characters). — G. Waterhouse, 1858: 26 (catalog of British species). — Schaum, 1859: 27 (catalog; European species). — LeConte, 1861a: 65 (characters). — Gredler, 1863: 108 (list of species from Tirol ). — LeConte, 1863: 24 (list of North American species). — Wencker and Silbermann, 1866: 30 (list of species; collecting notes; France ). — Gemminger and Harold, 1868: 606 (catalog). — Sharp, 1871d: 12 (list of British species). — Fauvel, 1873b: 86 [= 1873a: 372] (characters; key to species). — Redtenbacher, 1874: 221 (characters; key to Austrian species). — Fauvel, 1875a: XXIV [= 1875b: 226] (catalog). — Mulsant and Rey, 1877a: 237 (characters; key to French species). — Provancher, 1877: 251 (characters). — Heyden, 1880: 75 (list of species of Siberian region). — Duvivier, 1883: 159 (catalog). — Fowler, 1888: 294 (notes). — Seidlitz, 1891: 379 (key to some European species). — Seidlitz, 1891a: 404 (key to some European species). — Heyden, Reitter, and Weise, 1891: 105 (list of species of Europe and Caucasus). — Heyden, 1893: 43 (list of species of Siberian region). — Ganglbauer, 1895: 472 (characters; key to middle European species). — Heyden, 1896: 33 (list of species of Siberian region). — Casey, 1906: 424 (characters; key to species). — Heyden, Reitter, and Weise, 1906: 162 (list of species of Europe and Caucasus). — Reitter, 1908: 103, 107 (characters; key to species). — Reitter, 1909: 132, 133 (characters; key to species). Petri , 1912: 70 (list of species of Siebenbürgen). — Bernhauer and Schubert, 1914: 316 (world catalog; 9 species). — Johansen, 1914: 428 (characters; key to Danish species). — Leng, 1920: 106 (catalog of North American species). — Winkler, 1925: 373 (catalog for Palaearctic region). — Porta, 1926: 92 (characters; key to species of Italy ). — Portevin, 1929: 368 (key to French species). — Roubal, 1930: 364 (catalog; Slovakia ). — Joy, 1932: 127 (key to British species). — Scheerpeltz, 1933: 1320 (world catalog supplement). — Tottenham, 1945: 70 (synonym of Atrecus ; type species designation: Staphylinus pilicornis Paykull). — Tottenham, 1949: 370 (synonym of Atrecus ; type species: pilicornis ). — Blackwelder, 1952: 72 ( type species: pilicornis ). — V. Hansen, 1952: 7, 21 (characters; key to Danish species). — Coiffait, 1956a: 42 (key to French species). — Smetana, 1958: 117 (synonym of Atrecus ). — Palm, 1963b: 53 (characters; key to species of Sweden ). — Lohse, 1964: 167 (characters; key to central European species). — Scheerpeltz, 1968a: 63 (catalog of Austrian species). — Kasule, 1970: 74 (larval characters). Coiffait, 1972: 358 (characters; larval characters; type species: pilicornis ; key to species of western Palaearctic region; references) . — Pope, 1977: 28 (synonym of Atrecus ). — Uhlig, 1979: 252 (some species collected in Germany ). — Burakowski, Mroczkowski, and Stefańska, 1980: 22 (catalog; Poland ). — Smetana, 1982: 36 (synonym of Atrecus ). — Bordoni, 1982: 397 (characters; habitus; key to species of Italy ). — Tóth, 1989: 39 (characters; key to species of Hungary ). — Janák, 1993a: 47 (synonym of Atrecus ). affinis Paykull, 1789: 24 ( Staphylinus ; Type locality: Suecia ). — Erichson, 1839a: 422 ( Othius ; synonym of pilicornis ). — Erichson, 1839b: 297 ( Othius ; synonym of pilicornis ). — Jacquelin du Val, 1856: pl. 11, fig. 55 ( Atrecus ; [ Note: Misidentification: Bernhauer and Schubert, 1914: 316, cited pilicornis Jacquelin du Val as a synonym of affinis ; Jacquelin du Val did not describe the species as new, he attributed it to Paykull]; characters). — Kraatz, 1857d: 661 ( Baptolinus ; synonym of alternans ). — Fauvel, 1873b: 87 [= 1873a: 373] ( Baptolinus ; characters; France ; Scandinavia; Britain; Germany ; Switzerland ; Austria ; Italy ; Sicily ; Sardinia ; Caucasus). — Fauvel, 1874: 338 [= 1874b: 44] ( Baptolinus ; Spain ; Russia ). — Ganglbauer, 1895: 474 ( Baptolinus ; characters; Europe). — Reitter, 1908: 108 ( Baptolinus ; characters; Europe; Caucasus). — Reitter, 1909: 134 ( Baptolinus ; characters; Germany ). — Bernhauer and Schubert, 1914: 316 ( Baptolinus ; catalog). — Johansen, 1914: 429 ( Baptolinus ; characters; Denmark ). — Portevin, 1929: 368 ( Baptolinus ; characters; France ). — Scheerpeltz, 1933: 1320 ( Baptolinus ; catalog). — Paulian, 1941: 221 ( Baptolinus ; larval characters; France ). — V. Hansen, 1952: 22 ( Baptolinus ; characters; Denmark ). — Coiffait, 1956a: 43 ( Baptolinus ; characters in key). — Smetana, 1958: 121 ( Atrecus ; characters; ecology; Czechoslovakia ). — Palm, 1963b: 54 ( Baptolinus ; characters; Sweden ; Denmark ; Norway ; Finland ). — Lohse, 1964: 167 ( Baptolinus ; characters; central Europe). — Smetana, 1964d: 73 ( Atrecus ; Czechoslovakia ). — Coiffait, 1965e: 120 ( Othius ; Turkey ). — Horion, 1965: 103 ( Baptolinus ; Caucasus; Siberia; Finland ; Sweden ; Norway ; Denmark ; Britain; widespread in central and southern Europe; Spain ; Italy ; Dalmatia; Croatia ; Bosnia ; Albania ; Greece ). — Coiffait, 1966: 22 ( Baptolinus ; Turkey ). — Pototskaia, 1967: 49 ( Atrecus ; larval characters). — Szujecki, 1968a: 726 ( Atrecus ; cited as afiinis ; Poland ). — Bolov, 1969a: 328 ( Atrecus ; collecting notes; Russia ). — Coiffait, 1972: 363 ( Baptolinus ; characters; references; Europe; Scandinavia; Britain; Siberia; Turkey ). — Bordoni, 1973a: 192 ( Baptolinus ; Turkey ). — Bordoni, 1976: 236 ( Baptolinus ; Turkey ). — Pototskaia, 1976: 172 ( Atrecus ; larval characters). — Legner and Moore, 1977: 176 ( Atrecus ; found under bark). — Pope, 1977: 28 ( Atrecus ; Britain). — Bordoni, 1982: 404 ( Baptolinus ; characters; Italy ). — Bordoni, 1986b: 386 ( Baptolinus ; Slovakia ; Czechoslovakia ; Croatia ; Serbia ; Austria ; Macedonia ; Bulgaria ; Carniola). — Segers, 1986: 41 ( Baptolinus ; checklist of species; Belgium ). — Lucht, 1987: 103 ( Baptolinus ; checklist; Germany ; Poland ; Czechoslovakia ; Austria ; Switzerland ; France ; Benelux; Denmark ; Sweden ). — Tóth, 1989: 39 ( Baptolinus ; characters; Hungary ). — Ciceroni, 1993: 77 ( Atrecus ; Italy ). — Welch, 1993: 228 ( Atrecus ; ovariole number and ovary structure). — Janák, 1993a: 47 ( Atrecus ; Czech Republic ; Slovakia ). — Sparacio, 1995: 159 ( Atrecus ; characters; notes; Sicily ). — Ciceroni and Zanetti, 1995: 26 ( Atrecus ; Italy ). — Luff, 1996: 30 ( Atrecus ; collecting notes; Isle of Man ). — Assing, 1998c: 345, 350 ( Atrecus ; characters; discussion of variation; Georgia ; Turkey ; Greece ; Bulgaria ). — Owen, 1999b: 61 ( Atrecus ; captured underground at roots of old trees; Britain). — Fowles, Alexander, and Key, 1999: 129 ( Atrecus ; saproxylic species; status as endangered or threatened species; Britain). — Lott, 1999: 72 ( Atrecus ; saproxylic species; habitat; endangered status; Britain). — Owen, 2000: 251 ( Atrecus ; pitfall trap collections from underground at roots of trees; Britain). - DISTRIBUTION: Europe, Russia , Caucasus, Turkey . alternans Gravenhorst, 1802: 48 ( Staphylinus ; Type locality: Not cited, but presumably Brunsvigae ). — Latreille, 1804: 333 ( Staphylinus ; characters; Germany ). — Gravenhorst, 1806: 107 ( Staphylinus ). — Gyllenhal, 1810: 359 ( Staphylinus ; synonym of pilicornis Paykull ). — Curtis, 1829: 26 ( Xantholinus ; catalog; Britain). — C. Sahlberg, 1830: 335 ( Staphylinus ; synonym of pilicornis Paykull ). — Mannerheim, 1830: 34 ( Gyrohypnus ; Finland ). — Mannerheim, 1831: 448 ( Gyrohypnus ; Finland ). — Griffith and Pidgeon, 1832: 292 ( Xantholinus ). — Stephens, 1833: 254 ( Othius ; characters; England ). — Nordmann, 1837: 120 ( Gyrohypnus ; Germany ; Sweden ; Tauria). — Erichson, 1839a: 422 ( Othius ; synonym of pilicornis ). — Heer, 1839: 248 ( Othius ; characters; Switzerland ). — Erichson, 1839b: 297 ( Othius ; synonym of pilicornis ). — Stephens, 1839: 402 ( Othius ; characters; England ). — Laporte, 1840: 180 ( Gyrohypnus ; characters; France ). — Fairmaire and Laboulbène, 1856: 499 ( Othius ; synonym of pilicornis ). — Kraatz, 1857d: 660 ( Baptolinus ; characters; Germany ). — Redtenbacher, 1857: 982 ( Baptolinus ; characters; Austria ). — Thomson, 1860: 187 ( Gyrohypnus ; characters; Scandinavia). — Fauvel, 1865a: 298 [= 1865b: 48] ( Baptolinus ; habitat; France ). — Fauvel, 1867c: 114 [= 1868b: 213] ( Baptolinus ; synonym of affinis ). — Fauvel, 1873b: 87 [= 1873a: 373] ( Baptolinus ; synonym of affinis ). — Redtenbacher, 1874: 222 ( Baptolinus ; characters; Austria ). — Seidlitz, 1875: 281 ( Baptolinus ; characters; Baltic region). — J. Sahlberg, 1876: 33 ( Baptolinus ; Finland ). — Mulsant and Rey, 1877a: 243 ( Baptolinus ; characters; France ). — Fowler, 1888: 294 ( Baptolinus ; characters; Britain). — Ganglbauer, 1895: 474 ( Baptolinus ; synonym of affinis ). — Reitter, 1908: 108 ( Baptolinus ; synonym of affinis ). — Bernhauer and Schubert, 1914: 316 ( Baptolinus ; synonym of affinis ). — Scheerpeltz, 1933: 1321 ( Baptolinus ; synonym of affinis ). — Smetana, 1958: 121 ( Atrecus ; synonym of affinis ). — Coiffait, 1972: 363 ( Baptolinus ; synonym of affinis ). — Legner and Moore, 1977: 176 ( Atrecus ; found under bark). — Pope, 1977: 28 ( Atrecus ; synonym of affinis ). — Bordoni, 1982: 404 ( Baptolinus ; synonym of affinis ).