Contributions to the taxonomy, identification, and biogeography of Thaumatogelis Schwarz, 1995 (Hymenoptera: Ichneumonidae)
Author
Vas, Zoltán
Author
Schwarz, Martin
text
Zootaxa
2018
2018-07-11
4444
4
421
436
journal article
29694
10.11646/zootaxa.4444.4.4
7dfb506d-672d-40e4-862a-1d9e2b9bc564
1175-5326
1309859
F4E26BFC-A168-41CD-A0B0-E938CE0B5CAB
Thaumatogelis megaera
Vas sp. nov.
Material examined.
Holotype
: female,
Mongolia
,
Khövsgöl province
,
3 km
SW from
Somon Burenchaan
[on label:
Mongolia
,
Chövsgöl
aimak,
3 km
SW von Somon Burenchaan
],
1650 m
,
21.vi.–16.vii.1968
, Exp.
Dr. Z. Kaszab
1968,
Nr.
993,
pitfall trap
, leg.
Z. Kaszab
,
HNHM
Hym
. Coll. Id. No. 152917.—Paratypes: two females, Mongolia,
Dundgovi province
,
20 km
S from
Somon Delgertsogt
[on label:
Mongolia
,
Mittelgobi
aimak,
20 km
S von Somon Delgerzogt
],
1480 m
,
9.vi.1967
,
Exp. Dr. Z.
Kaszab 1967,
Nr.
779,
pitfall trap
, leg.
Z. Kaszab
,
HNHM
Hym
. Coll. Id. No. 152911–152912; female, Mongolia,
Khentii province
,
7 km
NE from
Somon Mörön
[on label:
Mongolia
,
Chentej
aimak,
7 km
NO von Somon Mörön
],
1200 m
,
28.vii.–21.viii.1965
,
Exp. Dr. Z.
Kaszab 1965,
Nr.
323,
pitfall trap
, leg.
Z. Kaszab
,
HNHM
Hym
. Coll. Id. No. 152913; female, Mongolia,
Khövsgöl province
,
Tunamal Lake
,
26 km
WSW from
Somon Sharga
[on label:
Mongolia
,
Chövsgöl
aimak, am
See Tunamal
nuur,
26 km
SW von Somon Scharga
],
1950 m
,
21.vi.–15.vii.1968
,
Exp. Dr. Z.
Kaszab 1968,
Nr.
996,
pitfall trap
, leg.
Z. Kaszab
,
HNHM
Hym
. Coll. Id. No. 152914; female, Mongolia,
Uvs province
,
3 km
WNW from SW corner of
Lake Orog
[on label:
Mongolia
,
Uva
aimak,
3 km
WNW von
SW
Ecke
des Sees Örög nuur],
1590 m
,
29.vi.1968
,
Exp. Dr. Z.
Kaszab 1968,
Nr.
1039, leg.
Z. Kaszab
,
HNHM
Hym
. Coll. Id. No. 152915; female, Mongolia,
Töv province
,
12 km
S from
Somon Bajanbaraat
[on label:
Mongolia
,
Central
aimak,
12 km
S von Somon Bajanbaraat
],
1380 m
,
8.vi.1967
,
Exp. Dr. Z.
Kaszab 1967,
Nr.
774,
pitfall trap
, leg.
Z. Kaszab
,
HNHM
Hym
. Coll. Id. No. 152916.—
The
holotype
and three
paratypes
(
HNHM
Hym. Coll. Id. No.
152911, 152914, 152915) are deposited in
HNHM
; one
paratype
(
HNHM
Hym. Coll. Id. No.
152913) is deposited in
BMNH
; one
paratype
(
HNHM
Hym. Coll. Id. No.
152912) is deposited in
OÖLM
; and one
paratype
(
HNHM
Hym.
Coll. Id. No.
152916) is deposited in
ZSM
.
Diagnosis.
T. megaera
Vas
sp. nov.
belongs to the morphological group of
Thaumatogelis
species characterised with enlarged sixth tergite. Within this group the newly described species can easily be separated from
T. vulpinus
(Gravenhorst)
,
T. neesii
(Förster)
,
T. robustus
(Seyrig)
,
T. pallens
Schwarz
, and from
T. femoralis
(Brischke)
by its distinctly longer, more slender first flagellomere (it is 2.6–3.0 × as long as wide in
T. megaera
Vas
sp. nov.
, while less than 2 × as long as wide in the other listed species).
T. asiaticus
Schwarz
differs from the newly described species by its strongly constricted head behind eyes (width of eye in dorsal view 1.8–2.1 × as long as gena in
T. asiaticus
Schwarz
, while 1.3–1.5 × as long as gena in
T. megaera
Vas
sp. nov.
).
T. inexspectatus
Schwarz
differs from the newly described species by its indistinctly punctate frons, entirely fused second and third tergites, shorter first flagellomere (2.0–2.5 × as long as wide in
T. inexspectatus
Schwarz
, while 2.6–3.0 × as long as wide in
T. megaera
Vas
sp. nov.
), and shorter hind tibia (3.7–3.8 × as long as high in
T. inexspectatus
Schwarz
, while 4.4–4.7 × as long as high in
T. megaera
Vas
sp. nov.
).
Description.
Female (
Figs 11–13
). Body length
4–5 mm
.
FIGURES 11–13.
Thaumatogelis megaera
Vas
sp. nov. 11
, holotype, dorsal habitus;
12
, holotype, lateral habitus;
13
, holotype, face in frontal view.
Head: Antenna filiform, not clavate, with 19–20 flagellomeres. First flagellomere 2.6–3.0 × as long as wide, fifth flagellomere 1.4–1.6 × as long as wide, first flagellomere 0.9–1.1 × as long as second flagellomere. Width of eye in dorsal view 1.3–1.5 × as long as gena, head behind eyes moderately constricted. OOL 0.6–0.7 × as long as POL. Inner eye orbits divergent ventrally. Vertex, gena and malar space strongly coriaceous and distinctly, densely punctate, punctures separated from each other usually by less than a puncture diameter. Frons strongly coriaceous, matt, and distinctly, densely punctate with punctures separated from each other by less than a puncture diameter, and with a few transverse wrinkles above toruli. Face strongly coriaceous, matt, and densely punctate with punctures separated from each other by usually less than a puncture diameter; face with short, dense, whitish hairs; face convex in lateral view. Clypeus less coriaceous than face, especially on apical half, but with stronger punctures; clypeus convex in profile, apical margin curved; hairs on clypeus longer than on face. Malar space 1.4–1.5 × as long as basal width of mandible. Lower mandibular tooth slightly shorter than upper tooth.
Mesosoma: Mesosoma coriaceous with dense but weak punctures, punctures separated from each other by usually less than a puncture diameter; mesosoma with dense, short, whitish hairs. Pronotum with horizontal wrinkles only at lower hind corner; pronotum and mesonotum fused with weak, indistinct margin. Mesonotum convex, coriaceous with dense but weak punctures; median furrow of mesonotum indistinct; scutellum very short, weakly separated; mesonotum 1.1–1.3 × as long as wide, and 1.5 × as long as anterior area of propodeum; groove between mesonotum and propodeum relatively shallow. Mesopleuron and metapleuron coriaceous with dense but weak punctures. Epicnemial carina ventrally and laterally strong, posterior transverse carina of mesosternum ventrally strong, mesosternum about 1.5–2.0 × as long as width of first flagellomere. Propodeum convex, about as high as or little higher than mesonotum in lateral view, anterior and posterior area indistinctly separated; anterior area coriaceous and densely punctate, punctures weak; posterior area coriaceous with dense but weak punctures and with a few, very weak irregular wrinkles; posterior transverse carina only laterally distinct. Legs coriaceous with dense but weak punctures; hind femur 4.4–4.7 × as long as high; tarsal claws simple, small, longer than arolium.
Metasoma: Metasoma with dense, short, whitish hairs. Dorsal surface of first tergite coriaceous with dense but weak punctures; first tergite stout, 1.3–1.5 × as long as apically wide. Second and third tergites dorsally not fused, though sometimes less distinctly separated, dorsal margin of second tergite always very distinct; second and third tergites laterally fused. Anterior two thirds of second tergite densely and strongly punctate, punctures separated from each other by less than a puncture diameter; on posterior third of second tergite punctures less dense, separated from each other usually by more than a puncture diameter. Third tergite densely and strongly punctate on anterior half, punctures separated from each other by less than a puncture diameter; posterior half of third tergite shinier and distinctly less densely punctate, punctures separated from each other by more than a puncture diameter. Sixth tergite enlarged, entirely covering seventh tergite. Ovipositor sheath relatively short, 0.3–0.5 × as long as hind tibia; ovipositor slender, dorsal margin without distinct, steep angular elevation before nodus, ventral margin moderately widened before nodus; ovipositor tip 4 × as long as high, with small notch; ventral margin of ovipositor tip distinctly narrowed, tapered toward apex and with very small teeth.
Colour: Head black with reddish brown patches at middle of face and at inner eye margins, sometimes reddish brown patches extend to lower gena, malar space and clypeus; mandible reddish brown, basal parts and teeth dark brown, other mouthparts yellowish brown. Scapus and pedicellus orange; basal flagellomeres orange, apical flagellomeres darker brownish. Mesosoma orange, darker brownish around sutures. Legs brown to reddish brown, femora usually darker than tibiae and tarsi, dorsoapical part of tibiae with more or less distinct yellowish brown patches. First tergite orange. Metasoma except first tergite blackish to dark brown, sometimes dark reddish brown at anterior and lateral parts of second tergite and at lateral part of third tergite. Posterior half of sixth tergite lighter brown. Ovipositor sheath brown.
Male: Unknown.
Distribution.
Currently known only from
Mongolia
.
Ecological note.
No host is known.
Etymology.
The specific epithet
megaera
is the Latinized name of one of the three Furies (or Erinyes) in the Greek and Roman mythology; proper noun in apposition, ending not to be changed.