<document id="EBF7A5202FF712C98150C3CD00246EBF" ID-ISSN="1175­5334" ID-ZooBank="6B088E9E-EAD4-4B8B-938B-178ADACF4CC3" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629809046025" checkinUser="felipe" docAuthor="Harzhauser, Mathias &amp; Landau, Bernard" docDate="2012" docId="387287FCC32BFFFFFF0DE06BE451F80C" docLanguage="en" docName="zt03472p071.pdf" docOrigin="Zootaxa 3472" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Trigonostoma Blainville 1827" docType="treatment" docVersion="2" lastPageNumber="55" masterDocId="C44BFF84C31DFFC9FF9AE57AE767FF93" masterDocTitle="A revision of the Neogene Cancellariid Gastropods of the Paratethys Sea 3472" masterLastPageNumber="71" masterPageNumber="1" pageNumber="55" updateTime="1699115141119" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="C99F385E169E932A27F20CEB5E910F9A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0F650F73904B466F2B136F41CA69B500">
<mods:title id="E6DB96C8C6549F1DD465F61E697DF039">A revision of the Neogene Cancellariid Gastropods of the Paratethys Sea 3472</mods:title>
</mods:titleInfo>
<mods:name id="EC9F642EB94DA196861E0A90D8D66FE5" type="personal">
<mods:role id="0C6C0FA12EB344834D821E33EC1052EE">
<mods:roleTerm id="41F8A0604D107285EBFEC5DC43C2DB63">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A0DEE3E81AFFD36D53B02FB89ADF7A97">Harzhauser, Mathias</mods:namePart>
</mods:name>
<mods:name id="452A729B86993B948785F2D3F0492C43" type="personal">
<mods:role id="B5501C08F4322025EA6250135F1C5A9D">
<mods:roleTerm id="CD86F31A9479B7694BF41F208D5E1190">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="83E838B8EB5B7C6A6D1CCBC916121B8B">Landau, Bernard</mods:namePart>
</mods:name>
<mods:typeOfResource id="B3DBF1880237A57A491AFACE38F1BE96">text</mods:typeOfResource>
<mods:relatedItem id="A67709EFB951BD742BE8A732673234A8" type="host">
<mods:titleInfo id="3D19896F8D87F7CD8165CB7EFA2AA65F">
<mods:title id="7A4A7D0BF0B807B95445F4837DB7639C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A49CEE4C04613B07A6E56D937DB351B0">
<mods:date id="663873111F5F59792B7F1047B2E10B0B">2012</mods:date>
<mods:detail id="1B31A04308E151B364482433C11EF896" type="pubDate">
<mods:number id="F7DEF0173BEC0D0BDF8B253575BEC144">2012-09-07</mods:number>
</mods:detail>
<mods:detail id="CFCF7A7AD804DBD616C5B9339A5BC869" type="volume">
<mods:number id="0D1534CE1CE0CBF473489873FD8B17EA">3472</mods:number>
</mods:detail>
<mods:extent id="D7034C177031E3C21DFFADE9483DFF5D" unit="page">
<mods:start id="7E949335F5ECABB58B3F3F2D93A7E5A2">1</mods:start>
<mods:end id="0613ECE0B49CA92F26DA7DDC791C4693">71</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="00CD0F6A18A16D6E1FCD7EA7821C9ACA">journal article</mods:classification>
<mods:identifier id="16CEFE876A7BA3981644345C7486F3AE" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="2FFB1DA57569193A59CCB799F3B96A2D" type="ZooBank">6B088E9E-EAD4-4B8B-938B-178ADACF4CC3</mods:identifier>
</mods:mods>
<treatment id="387287FCC32BFFFFFF0DE06BE451F80C" LSID="urn:lsid:plazi:treatment:387287FCC32BFFFFFF0DE06BE451F80C" httpUri="http://treatment.plazi.org/id/387287FCC32BFFFFFF0DE06BE451F80C" lastPageNumber="55" pageId="54" pageNumber="55">
<subSubSection id="F8C16561C32BFFFFFF0DE06BE5CBFAB9" box="[151,684,1296,1323]" pageId="54" pageNumber="55" type="nomenclature">
<paragraph id="B06436EAC32BFFFFFF0DE06BE5CBFAB9" blockId="54.[151,684,1296,1323]" box="[151,684,1296,1323]" pageId="54" pageNumber="55">
<heading id="EB2C8186C32BFFFFFF0DE06BE5CBFAB9" bold="true" box="[151,684,1296,1323]" fontSize="11" level="1" pageId="54" pageNumber="55" reason="1">
<emphasis id="82AFEAF8C32BFFFFFF0DE06BE5CBFAB9" bold="true" box="[151,684,1296,1323]" pageId="54" pageNumber="55">
<taxonomicName id="77DB4D69C32BFFFFFF0DE06BE627FAB8" ID-CoL="7PJWP" authorityName="Blainville" authorityYear="1827" box="[151,320,1297,1323]" class="Gastropoda" family="Cancellariidae" genus="Trigonostoma" kingdom="Animalia" order="Neogastropoda" pageId="54" pageNumber="55" phylum="Mollusca" rank="genus">
<emphasis id="82AFEAF8C32BFFFFFF0DE06BE627FAB8" bold="true" box="[151,320,1297,1323]" italics="true" pageId="54" pageNumber="55">Trigonostoma</emphasis>
</taxonomicName>
? 
<taxonomicName id="77DB4D69C32BFFFFFEC6E06BE5CBFAB9" authority="(Cossmann, 1915)" baseAuthorityName="Cossmann" baseAuthorityYear="1915" box="[348,684,1296,1323]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="boettgeri">
<emphasis id="82AFEAF8C32BFFFFFEC6E06BE6A1FAB8" bold="true" box="[348,454,1297,1323]" italics="true" pageId="54" pageNumber="55">boettgeri</emphasis>
(Cossmann, 1915)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F8C16561C32BFFFFFF0DE022E42BFA38" pageId="54" pageNumber="55" type="reference_group">
<paragraph id="B06436EAC32BFFFFFF0DE022E385FAFD" blockId="54.[151,1250,1368,1451]" box="[151,1250,1368,1390]" pageId="54" pageNumber="55">
<treatmentCitationGroup id="90CB11C4C32BFFFFFF0DE022E385FAFD" box="[151,1250,1368,1390]" pageId="54" pageNumber="55">
<treatmentCitation id="317A10FBC32BFFFFFF0DE022E469FAFD" author="Boettger, O." box="[151,782,1368,1390]" page="138" pageId="54" pageNumber="55" year="1906">
<taxonomicName id="77DB4D69C32BFFFFFF0DE022E469FAFD" ID-CoL="6CYH8" authority="Boettger, 1906: 138" authorityName="Boettger" authorityPageNumber="138" authorityYear="1906" box="[151,782,1368,1390]" class="Gastropoda" family="Discohelicidae" genus="Discohelix" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="calculiformis" subGenus="Pseudomalaxis">
<emphasis id="82AFEAF8C32BFFFFFF0DE022E664FAFD" box="[151,259,1368,1390]" italics="true" pageId="54" pageNumber="55">Discohelix</emphasis>
(
<emphasis id="82AFEAF8C32BFFFFFE88E022E6CCFAFD" box="[274,427,1368,1390]" italics="true" pageId="54" pageNumber="55">Pseudomalaxis</emphasis>
) 
<emphasis id="82AFEAF8C32BFFFFFE20E022E558FAFD" box="[442,575,1368,1390]" italics="true" pageId="54" pageNumber="55">calculiformis</emphasis>
<bibRefCitation id="D44A4B1BC32BFFFFFDDCE022E469FAFD" author="Boettger, O." box="[582,782,1368,1390]" pageId="54" pageNumber="55" pagination="1 - 99" refId="ref56572" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso - Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99." type="journal article" year="1906">Boettger, 1906: 138</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[non 
<taxonomicName id="77DB4D69C32BFFFFFCD0E022E3B0FAFD" ID-CoL="6CYH8" authority="Dunker, 1847" authorityName="Dunker" authorityYear="1847" box="[842,1239,1368,1390]" class="Gastropoda" family="Discohelicidae" genus="Discohelix" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="calculiformis">
<emphasis id="82AFEAF8C32BFFFFFCD0E022E326FAFD" box="[842,1089,1368,1390]" italics="true" pageId="54" pageNumber="55">Discohelix calculiformis</emphasis>
<bibRefCitation id="D44A4B1BC32BFFFFFBD2E022E3B0FAFD" author="Dunker, W." box="[1096,1239,1368,1390]" pageId="54" pageNumber="55" pagination="128 - 133" refId="ref57908" refString="Dunker, W. (1847) Ueber einige neue Versteinerungen aus verschiedenen Gebirgsformationen. Palaeontographica, 1, 128 - 133." type="journal article" year="1847">Dunker, 1847</bibRefCitation>
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="B06436EAC32BFFFFFF0DE00DE452FA1F" blockId="54.[151,1250,1368,1451]" box="[151,821,1398,1421]" pageId="54" pageNumber="55">
<taxonomicName id="77DB4D69C32BFFFFFF0DE00DE511FA1F" authority="Cossmann, 1915: 143" authorityName="Cossmann" authorityPageNumber="143" authorityYear="1915" box="[151,630,1398,1421]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="boettgeri">
<emphasis id="82AFEAF8C32BFFFFFF0DE00DE6F5FA1E" box="[151,402,1399,1421]" italics="true" pageId="54" pageNumber="55">Pseudomalaxis boettgeri</emphasis>
Cossmann, 1915: 143
</taxonomicName>
, pl. 12, figs 22-24.
</paragraph>
<paragraph id="B06436EAC32BFFFFFF0DE0EFE42BFA38" blockId="54.[151,1250,1368,1451]" box="[151,844,1429,1451]" pageId="54" pageNumber="55">
<treatmentCitationGroup id="90CB11C4C32BFFFFFF0DE0EFE42BFA38" box="[151,844,1429,1451]" pageId="54" pageNumber="55">
<taxonomicName id="77DB4D69C32BFFFFFF0DE0EFE566FA38" authority="Cossmann" authorityName="Cossmann" authorityYear="1915" box="[151,513,1429,1451]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="boettgeri">
<emphasis id="82AFEAF8C32BFFFFFF0DE0EFE6F5FA38" box="[151,402,1429,1451]" italics="true" pageId="54" pageNumber="55">Pseudomalaxis boettgeri</emphasis>
Cossmann
</taxonomicName>
— 
<treatmentCitation id="317A10FBC32BFFFFFD81E0EFE5D8FA38" author="Zilch, A." box="[539,703,1429,1451]" page="219" pageId="54" pageNumber="55" year="1934">
<bibRefCitation id="D44A4B1BC32BFFFFFD81E0EFE5D8FA38" author="Zilch, A." box="[539,703,1429,1451]" pageId="54" pageNumber="55" pagination="193 - 302" refId="ref63428" refString="Zilch, A. (1934) Zur Fauna des Mittel - Miocans von Kostej (Banat). Typus-Bestimmung und Tafeln zu O. BOETTGER's Bearbeitungen. Senckenbergiana, 16, 193 - 302." type="journal article" year="1934">Zilch 1934: 219</bibRefCitation>
</treatmentCitation>
, pl. 7, fig. 22.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F8C16561C32BFFFFFF0DE0A2E66DF987" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="B06436EAC32BFFFFFF0DE0A2E66DF987" blockId="54.[151,1436,1495,1951]" pageId="54" pageNumber="55">
<emphasis id="82AFEAF8C32BFFFFFF0DE0A2E623FA63" bold="true" box="[151,324,1495,1520]" inLineHeading="true" pageId="54" pageNumber="55" reason="1">Type material.</emphasis>
<materialsCitation id="00B33CB7C32BFFFFFED1E0A2E537FA7C" box="[331,592,1495,1520]" collectingDate="2375-12" collectionCode="SMF" pageId="54" pageNumber="55" specimenCount="1" typeStatus="syntype">
<typeStatus id="6F608848C32BFFFFFED1E0A2E6C5FA63" box="[331,418,1496,1520]" pageId="54" pageNumber="55" type="syntype">syntype</typeStatus>
<collectionCode id="D6CAAE2FC32BFFFFFE33E0ADE686FA7C" box="[425,481,1495,1519]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="54" pageNumber="55">SMF</collectionCode>
<date id="C465102AC32BFFFFFE7DE0A2E537FA7C" box="[487,592,1495,1519]" pageId="54" pageNumber="55" value="2375-12">
<collectingDate id="D421E9C2C32BFFFFFE7DE0A2E537FA7C" box="[487,592,1495,1519]" pageId="54" pageNumber="55" value="2375-12">XII 2375</collectingDate>
</date>
</materialsCitation>
a, 
<materialsCitation id="00B33CB7C32BFFFFFDFDE0ADE661F987" collectionCode="SMF" collectorName="de Sus" country="Germany" location="CoSteiu de Sus" municipality="Middle Miocene" pageId="54" pageNumber="55" specimenCount="1" typeStatus="syntype">
<collectionCode id="D6CAAE2FC32BFFFFFDFDE0ADE5F8FA7C" box="[615,671,1495,1519]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="54" pageNumber="55">SMF</collectionCode>
Frankfurt (
<collectingCountry id="C8CC767AC32BFFFFFC84E0ADE4EBFA7C" box="[798,908,1495,1519]" name="Germany" pageId="54" pageNumber="55">Germany</collectingCountry>
), 
<location id="B5046031C32BFFFFFC05E0ADE322FA7C" LSID="urn:lsid:plazi:treatment:387287FCC32BFFFFFF0DE06BE451F80C:B5046031C32BFFFFFC05E0ADE322FA7C" box="[927,1093,1495,1519]" country="Germany" municipality="Middle Miocene" name="CoSteiu de Sus" pageId="54" pageNumber="55">
CoŞteiu 
<collectorName id="1D2E533CC32BFFFFFC61E0ADE322FA7C" box="[1019,1093,1495,1519]" pageId="54" pageNumber="55">de Sus</collectorName>
</location>
(
<collectingCountry id="C8CC767AC32BFFFFFBC9E0ADE3D9FA7C" box="[1107,1214,1495,1519]" name="Romania" pageId="54" pageNumber="55">Romania</collectingCountry>
), Badenian (
<collectingMunicipality id="5000AC90C32BFFFFFAD3E0ADE799F987" pageId="54" pageNumber="55">Middle Miocene</collectingMunicipality>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8C16561C32BFFFFFF59E365E41EF89C" pageId="54" pageNumber="55" type="discussion">
<paragraph id="B06436EAC32BFFFFFF59E365E41EF89C" blockId="54.[151,1436,1495,1951]" pageId="54" pageNumber="55">
<emphasis id="82AFEAF8C32BFFFFFF59E365E62FF9AB" bold="true" box="[195,328,1567,1592]" inLineHeading="true" pageId="54" pageNumber="55" reason="1">Discussion.</emphasis>
<bibRefCitation id="D44A4B1BC32BFFFFFEC8E35AE560F9A4" author="Boettger, O." box="[338,519,1567,1592]" pageId="54" pageNumber="55" pagination="1 - 99" refId="ref56572" refString="Boettger, O. (1906) Zur Kenntnis der Fauna der mittelmiocanen Schichten von Kostej im Krasso - Szorenyer Komitat. Gasteropoden und Anneliden III. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften zu Hermannstadt, 54, 1 - 99." type="journal article" year="1906">Boettger (1906)</bibRefCitation>
described a juvenile cancellariid from the early Badenian of the Transylvanian Basin as 
<taxonomicName id="77DB4D69C32BFFFFFF67E33EE576F9CF" authorityName="Dunker" authorityYear="1847" box="[253,529,1604,1628]" class="Gastropoda" family="Discohelicidae" genus="Discohelix" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="calculiformis">
<emphasis id="82AFEAF8C32BFFFFFF67E33EE576F9CF" box="[253,529,1604,1628]" italics="true" pageId="54" pageNumber="55">Discohelix calculiformis</emphasis>
</taxonomicName>
. This name was preoccupied for a Jurassic species described by 
<bibRefCitation id="D44A4B1BC32BFFFFFB75E33EE2FFF9CF" author="Dunker, W." box="[1263,1432,1604,1628]" pageId="54" pageNumber="55" pagination="128 - 133" refId="ref57908" refString="Dunker, W. (1847) Ueber einige neue Versteinerungen aus verschiedenen Gebirgsformationen. Palaeontographica, 1, 128 - 133." type="journal article" year="1847">Dunker (1847)</bibRefCitation>
. Therefore, Cossmann (1915) proposed the replacement name 
<taxonomicName id="77DB4D69C32BFFFFFCD1E312E307F913" authorityName="Cossmann" authorityYear="1915" box="[843,1120,1640,1664]" class="Gastropoda" family="Architectonicidae" genus="Pseudomalaxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="boettgeri">
<emphasis id="82AFEAF8C32BFFFFFCD1E312E307F913" box="[843,1120,1640,1664]" italics="true" pageId="54" pageNumber="55">Pseudomalaxis boettgeri</emphasis>
</taxonomicName>
. The presence of faint columellar folds excludes a placement in the 
<taxonomicName id="77DB4D69C32BFFFFFDFFE3F6E44BF937" authorityName="J.E.Gray" authorityYear="1850" box="[613,812,1676,1700]" class="Gastropoda" family="Architectonicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="54" pageNumber="55" phylum="Mollusca" rank="family">Architectonicidae</taxonomicName>
and consequently 
<bibRefCitation id="D44A4B1BC32BFFFFFC65E3F6E253F937" author="Petit, R. E. &amp; Harasewych, M. G." box="[1023,1332,1676,1700]" pageId="54" pageNumber="55" pagination="3 - 161" refId="ref61369" refString="Petit, R. E. &amp; Harasewych, M. G. (2005) Catalogue of the superfamily Cancellarioidea Forbes and Hanley, 1851 (Gastropoda: Prosobranchia) 2 nd edition. Zootaxa, 1102, 3 - 161." type="journal article" year="2005">Petit &amp; Harasewych (2005)</bibRefCitation>
included it in 
<taxonomicName id="77DB4D69C32BFFFFFF51E3CAE601F954" authorityName="Blainville" authorityYear="1827" box="[203,358,1712,1735]" class="Gastropoda" family="Cancellariidae" genus="Trigonostoma" kingdom="Animalia" order="Neogastropoda" pageId="54" pageNumber="55" phylum="Mollusca" rank="genus" sensu="lato">
<emphasis id="82AFEAF8C32BFFFFFF51E3CAE601F954" box="[203,358,1712,1735]" italics="true" pageId="54" pageNumber="55">Trigonostoma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="999C5783C32BFFFFFEF7E3D5E6E9F954" box="[365,398,1711,1735]" pageId="54" pageNumber="55" sensu="lato">s.l.</taxonomicNameLabel>
The available material does not allow clarifying the status of this species. The characteristic planorbiform early teleoconch is unlike that of any other known species and therefore we consider it a valid species, despite the very juvenile and fragmentary material available.
</paragraph>
</subSubSection>
<subSubSection id="F8C16561C32BFFFFFF59E261E451F80C" pageId="54" pageNumber="55" type="distribution">
<paragraph id="B06436EAC32BFFFFFF59E261E55AF8C4" blockId="54.[151,1436,1495,1951]" pageId="54" pageNumber="55">
<emphasis id="82AFEAF8C32BFFFFFF59E261E63BF8A7" bold="true" box="[195,348,1819,1844]" inLineHeading="true" pageId="54" pageNumber="55" reason="1">Distribution.</emphasis>
A rare species, which is known so far only from the early Badenian of the Transylvanian Basin and the North Alpine Foreland Basin.
</paragraph>
<paragraph id="B06436EAC32BFFFFFF59E21EE451F80C" blockId="54.[151,1436,1495,1951]" pageId="54" pageNumber="55">
<emphasis id="82AFEAF8C32BFFFFFF59E21EE65BF8EF" box="[195,316,1892,1916]" italics="true" pageId="54" pageNumber="55">Paratethys</emphasis>
—Badenian: Transylvanian Basin (
<collectingCountry id="C8CC767AC32BFFFFFD51E21EE451F8EF" box="[715,822,1892,1916]" name="Romania" pageId="54" pageNumber="55">Romania</collectingCountry>
: CoŞteiu de Sus), North Alpine Foreland Basin (
<collectingCountry id="C8CC767AC32BFFFFFAFEE21EE7A7F80C" name="Austria" pageId="54" pageNumber="55">Austria</collectingCountry>
: Grund; private collection of A. Breitenberger, 
<collectingCountry id="C8CC767AC32BFFFFFD48E2FDE44DF80C" box="[722,810,1927,1951]" name="Austria" pageId="54" pageNumber="55">Austria</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>