<document id="4134FFC71937ECAF0568EBD8DA549AE1" ID-DOI="10.11606/1807-0205/2024.64.031" ID-ISSN="1807-0205" ID-Zenodo-Dep="14705562" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737339390010" checkinUser="felipe" docAuthor="Simone, Luiz Ricardo L." docDate="2024" docId="4008878EFFA9A8448943DFF6FA3FFCE1" docLanguage="en" docName="PapAvZool.64.e202464031.pdf" docOrigin="Papéis Avulsos de Zoologia 64" docSource="https://doi.org/10.11606/1807-0205/2024.64.031" docStyle="DocumentStyle:6844D5473582683DD13A9A75B001759F.4:PapAvZool.2018-.journal_article" docStyleId="6844D5473582683DD13A9A75B001759F" docStyleName="PapAvZool.2018-.journal_article" docStyleVersion="4" docTitle="Scaphopoda Bronn 1862" docType="treatment" docVersion="2" lastPageNumber="97" masterDocId="BC31FFF6FFC9A8248835DF2CFFA6FFD2" masterDocTitle="New species, misidentifications and problematic taxonomy of some Atlantic South American marine mollusks: a review" masterLastPageNumber="104" masterPageNumber="1" pageNumber="97" updateTime="1737393675899" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0" zenodo-license-figures="CC-BY-NC-4.0">
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<mods:title id="39C6CD40BFD260FF241EAE5049EDA7C2">New species, misidentifications and problematic taxonomy of some Atlantic South American marine mollusks: a review</mods:title>
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<mods:namePart id="18FF209E9AF7F087F15D2BA5C3199E6F">Simone, Luiz Ricardo L.</mods:namePart>
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<mods:title id="64D67CDE82351C44B767C5C7114073A9">Papéis Avulsos de Zoologia</mods:title>
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<mods:date id="EC5D4F1E004192CE132061D53E94DD97">2024</mods:date>
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<mods:number id="B42F9AC50723F511875997A46DE0C558">64</mods:number>
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<mods:classification id="68CBEC483CA1C9932EB06E2B1A467B83">journal article</mods:classification>
<mods:identifier id="2DF2AEC405EE05B733591C630EE41BD1" type="DOI">10.11606/1807-0205/2024.64.031</mods:identifier>
<mods:identifier id="99A7AA20B8436A373C549D370E7682C7" type="ISSN">1807-0205</mods:identifier>
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<treatment id="4008878EFFA9A8448943DFF6FA3FFCE1" LSID="urn:lsid:plazi:treatment:4008878EFFA9A8448943DFF6FA3FFCE1" httpUri="http://treatment.plazi.org/id/4008878EFFA9A8448943DFF6FA3FFCE1" lastPageNumber="97" pageId="96" pageNumber="97">
<subSubSection id="80BB6513FFA9A8448943DFF6FD6DFE81" pageId="96" pageNumber="97" type="nomenclature">
<paragraph id="C81E3698FFA9A8448943DFF6FDE3FEC1" blockId="96.[184,780,218,435]" pageId="96" pageNumber="97">
<heading id="935681F4FFA9A8448943DFF6FDE3FEC1" centered="true" fontSize="10" level="2" pageId="96" pageNumber="97" reason="2">
<emphasis id="FAD5EA8AFFA9A8448943DFF6FDE3FEC1" bold="true" pageId="96" pageNumber="97">
Class 
<taxonomicName id="0FA14D1BFFA9A844898DDFF6FDEBFF21" ID-CoL="8ZXV9" authorityName="Bronn" authorityYear="1862" box="[440,589,218,243]" class="Scaphopoda" kingdom="Animalia" pageId="96" pageNumber="97" phylum="Mollusca" rank="class">Scaphopoda</taxonomicName>
Order 
<taxonomicName id="0FA14D1BFFA9A84489FCDFD6FDE3FEC1" authorityName="Starobogatov" authorityYear="1974" box="[457,581,250,275]" class="Scaphopoda" kingdom="Animalia" order="Dentaliida" pageId="96" pageNumber="97" phylum="Mollusca" rank="order">Dentaliida</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="C81E3698FFA9A8448946DE36FD6DFE81" blockId="96.[184,780,218,435]" pageId="96" pageNumber="97">
<emphasis id="FAD5EA8AFFA9A8448946DE36FD6DFE81" bold="true" pageId="96" pageNumber="97">
Family 
<taxonomicName id="0FA14D1BFFA9A84489F2DE36FDF7FEE1" authorityName="Children" authorityYear="1834" box="[455,593,282,307]" class="Scaphopoda" family="Dentaliidae" kingdom="Animalia" order="Dentaliida" pageId="96" pageNumber="97" phylum="Mollusca" rank="family">Dentaliidae</taxonomicName>
Genus 
<taxonomicName id="0FA14D1BFFA9A844897CDE16FD6DFE81" authority="Pilsbry &amp; Sharp, 1897" authorityName="Pilsbry &amp; Sharp" authorityYear="1897" box="[329,715,314,339]" class="Scaphopoda" family="Dentaliidae" genus="Graptacme" kingdom="Animalia" order="Dentaliida" pageId="96" pageNumber="97" phylum="Mollusca" rank="genus">
<emphasis id="FAD5EA8AFFA9A844897CDE16FE6CFE81" bold="true" box="[329,458,314,339]" italics="true" pageId="96" pageNumber="97">Graptacme</emphasis>
Pilsbry &amp; Sharp, 1897
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="80BB6513FFA9A844888DDE76FDBEFDC1" pageId="96" pageNumber="97" type="description">
<paragraph id="C81E3698FFA9A844888DDE76FCADFE61" blockId="96.[184,780,218,435]" pageId="96" pageNumber="97">
<emphasis id="FAD5EA8AFFA9A844888DDE76FCADFE61" bold="true" pageId="96" pageNumber="97">
<taxonomicName id="0FA14D1BFFA9A844888DDE76FDD2FEA1" authority="Fogarolli &amp; Simone" authorityName="Fogarolli &amp; Simone" authorityYear="2024" box="[184,628,346,371]" class="Scaphopoda" family="Dentaliidae" genus="Graptacme" kingdom="Animalia" order="Dentaliida" pageId="96" pageNumber="97" phylum="Mollusca" rank="species" species="obtura" status="sp. nov.">
<emphasis id="FAD5EA8AFFA9A844888DDE76FE2AFEA1" bold="true" box="[184,396,346,371]" italics="true" pageId="96" pageNumber="97">Graptacme obtura</emphasis>
Fogarolli &amp; Simone
</taxonomicName>
, 
<taxonomicNameLabel id="E1E657F1FFA9A8448A4BDE76FCAAFEA1" box="[638,780,346,371]" pageId="96" pageNumber="97" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="509A2A1DFFA9A8448993DE56FE55FE41" box="[422,499,378,403]" captionStart="Figure 64" captionStartId="96.[170,229,2042,2067]" captionTargetBox="[170,1493,859,2012]" captionTargetId="figure-228@96.[170,1493,859,2012]" captionTargetPageId="96" captionText="Figure 64. Shell of types of Graptacme obtura and G. semistriolata: (A) G. obtura holotype MZSP 166760 (L 35.6 mm), dorsal view; (B) same, right view; (C) same, detail of posterior region showing slit; (D-F) G. obtura paratype MZSP 121837 (L 36.4 mm), same views as preceding specimen; (G-I) G. semistriolata MZSP 47357 (L 26.4 mm),same views as first specimen.Scales:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14705714" httpUri="https://zenodo.org/record/14705714/files/figure.png" pageId="96" pageNumber="97">Fig. 64</figureCitation>
A-F) https://zoobank.org/ 
<uuid id="BC070C4DFFA9A844894FDEB6FCADFE61" box="[378,779,410,435]" pageId="96" pageNumber="97">41CB788D-9EA1-4011-8DD2-3354945D0AC4</uuid>
</emphasis>
</paragraph>
<paragraph id="C81E3698FFA9A844889FDEF6FDBEFDC1" blockId="96.[170,794,474,531]" pageId="96" pageNumber="97">
<emphasis id="FAD5EA8AFFA9A844889FDEF6FF50FE21" bold="true" box="[170,246,474,499]" pageId="96" pageNumber="97">Types:</emphasis>
<materialsCitation id="78C93CC5FFA9A8448935DEF6FDF7FE21" box="[256,593,474,499]" collectionCode="MZSP" pageId="96" pageNumber="97" specimenCode="MZSP 166760" specimenCount="1" typeStatus="holotype">
<typeStatus id="171A883AFFA9A8448935DEF6FEC1FE21" box="[256,359,474,499]" pageId="96" pageNumber="97" type="holotype">Holotype</typeStatus>
<specimenCode id="98079EE3FFA9A8448947DEF6FDB7FE21" box="[370,529,474,499]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="96" pageNumber="97" type="Museum">MZSP 166760</specimenCode>
, spm
</materialsCitation>
. 
<typeStatus id="171A883AFFA9A8448A6BDEF6FD6BFE21" box="[606,717,474,499]" pageId="96" pageNumber="97" type="paratype">Paratypes</typeStatus>
: MZSP 121837, 6 spm from type locality.
</paragraph>
</subSubSection>
<subSubSection id="80BB6513FFA9A844889FDD16FD44FD41" pageId="96" pageNumber="97" type="materials_examined">
<paragraph id="C81E3698FFA9A844889FDD16FD44FD41" blockId="96.[170,794,570,659]" pageId="96" pageNumber="97">
<materialsCitation id="78C93CC5FFA9A844889FDD16FDF6FD81" box="[170,592,570,595]" country="Brazil" location="Espirito Santo" pageId="96" pageNumber="97" specimenCount="1" stateProvince="Espirito Santo" typeStatus="holotype">
<emphasis id="FAD5EA8AFFA9A844889FDD16FEEEFD81" bold="true" box="[170,328,570,595]" pageId="96" pageNumber="97">
<typeStatus id="171A883AFFA9A844889FDD16FF45FD81" box="[170,227,570,595]" pageId="96" pageNumber="97">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="B0B67608FFA9A844897ADD16FE07FD81" box="[335,417,570,595]" name="Brazil" pageId="96" pageNumber="97">BRAZIL</collectingCountry>
. 
<emphasis id="FAD5EA8AFFA9A844899CDD16FDF6FD81" bold="true" box="[425,592,570,595]" pageId="96" pageNumber="97">
<collectingRegion id="0A65F87AFFA9A844899CDD16FDF6FD81" box="[425,592,570,595]" country="Brazil" name="Espirito Santo" pageId="96" pageNumber="97">Espírito Santo</collectingRegion>
</emphasis>
</materialsCitation>
; Trindade oceanic island, Calheta beach, 
<geoCoordinate id="AD95505FFFA9A844899FDD76FD93FDA1" box="[426,565,601,627]" degrees="20" direction="south" minutes="30" orientation="latitude" pageId="96" pageNumber="97" precision="1" seconds="37.6" value="-20.510445">20°30′37.6″S</geoCoordinate>
<geoCoordinate id="AD95505FFFA9A8448A0BDD76FD72FDA1" box="[574,724,601,627]" degrees="29" direction="west" minutes="18" orientation="longitude" pageId="96" pageNumber="97" precision="1" seconds="28.1" value="-29.307804">29°18′28.1″W</geoCoordinate>
, 
<quantity id="0F599B7DFFA9A8448AEADD76FCBFFDA1" box="[735,793,602,627]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="96" pageNumber="97" unit="m" value="17.0">17 m</quantity>
[Protrindade Project, J.B. MendonÇa col., 
<date id="BC1F1058FFA9A8448A5FDD56FD7FFD41" box="[618,729,634,659]" pageId="96" pageNumber="97" value="2014-10-26">26.x.2014</date>
].
</paragraph>
</subSubSection>
<subSubSection id="80BB6513FFA9A844889FDD96FE74FCE1" pageId="96" pageNumber="97" type="diagnosis">
<paragraph id="C81E3698FFA9A844889FDD96FE74FCE1" blockId="96.[170,794,698,819]" pageId="96" pageNumber="97">
<emphasis id="FAD5EA8AFFA9A844889FDD96FE80FD01" bold="true" box="[170,294,698,723]" pageId="96" pageNumber="97">Diagnosis:</emphasis>
Species from Trindade oceanic island with striae and slit in posterior region longer than 11% of shell length. Rate of increase 
<quantity id="0F599B7DFFA9A8448981DDD6FD85FCC1" box="[436,547,762,787]" metricMagnitude="-2" metricUnit="m" metricValue="7.8" pageId="96" pageNumber="97" unit="mm" value="78.0">0,078 mm</quantity>
/mm of length. Strong bilabial posterior aperture.
</paragraph>
</subSubSection>
<subSubSection id="80BB6513FFA9A8448B50DFF6FB94FDA1" pageId="96" pageNumber="97" type="description">
<paragraph id="C81E3698FFA9A8448B50DFF6FB94FDA1" blockId="96.[869,1493,218,627]" pageId="96" pageNumber="97">
<emphasis id="FAD5EA8AFFA9A8448B50DFF6FC5FFF21" bold="true" box="[869,1017,218,243]" pageId="96" pageNumber="97">Description:</emphasis>
Size about 
<quantity id="0F599B7DFFA9A8448CACDFF6FB52FF21" box="[1177,1268,218,243]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="96" pageNumber="97" unit="mm" value="36.0">36 mm</quantity>
; walls thin, translucid, color white; well curved, section circular (
<figureCitation id="509A2A1DFFA9A8448B59DE36FB96FEE1" box="[876,1072,282,307]" captionStart="Figure 64" captionStartId="96.[170,229,2042,2067]" captionTargetBox="[170,1493,859,2012]" captionTargetId="figure-228@96.[170,1493,859,2012]" captionTargetPageId="96" captionText="Figure 64. Shell of types of Graptacme obtura and G. semistriolata: (A) G. obtura holotype MZSP 166760 (L 35.6 mm), dorsal view; (B) same, right view; (C) same, detail of posterior region showing slit; (D-F) G. obtura paratype MZSP 121837 (L 36.4 mm), same views as preceding specimen; (G-I) G. semistriolata MZSP 47357 (L 26.4 mm),same views as first specimen.Scales:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14705714" httpUri="https://zenodo.org/record/14705714/files/figure.png" pageId="96" pageNumber="97">Fig. 64A, B, D, E</figureCitation>
). widening smooth (tax of increase about 
<quantity id="0F599B7DFFA9A8448B9BDE16FBB9FE81" box="[942,1055,314,339]" metricMagnitude="-5" metricUnit="m" metricValue="7.8" pageId="96" pageNumber="97" unit="mm" value="0.078">0.078 mm</quantity>
/mm of length). Sculpture absent, shell smooth, glossy; except for posterior 11-15% of shell length possessing longitudinal, delicate striae in entire circumference; striae gradually disappearing anteriorly (
<figureCitation id="509A2A1DFFA9A8448B59DE96FC44FE01" box="[876,994,442,467]" captionStart="Figure 64" captionStartId="96.[170,229,2042,2067]" captionTargetBox="[170,1493,859,2012]" captionTargetId="figure-228@96.[170,1493,859,2012]" captionTargetPageId="96" captionText="Figure 64. Shell of types of Graptacme obtura and G. semistriolata: (A) G. obtura holotype MZSP 166760 (L 35.6 mm), dorsal view; (B) same, right view; (C) same, detail of posterior region showing slit; (D-F) G. obtura paratype MZSP 121837 (L 36.4 mm), same views as preceding specimen; (G-I) G. semistriolata MZSP 47357 (L 26.4 mm),same views as first specimen.Scales:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14705714" httpUri="https://zenodo.org/record/14705714/files/figure.png" pageId="96" pageNumber="97">Fig. 64C, F</figureCitation>
). Anterior aperture rounded; peristome simple, with cutting edges. Posterior aperture with very narrow, elongated slit in ventral, sometimes in lateral surface (
<figureCitation id="509A2A1DFFA9A8448BD7DD36FBFAFDE1" box="[994,1116,538,563]" captionStart="Figure 64" captionStartId="96.[170,229,2042,2067]" captionTargetBox="[170,1493,859,2012]" captionTargetId="figure-228@96.[170,1493,859,2012]" captionTargetPageId="96" captionText="Figure 64. Shell of types of Graptacme obtura and G. semistriolata: (A) G. obtura holotype MZSP 166760 (L 35.6 mm), dorsal view; (B) same, right view; (C) same, detail of posterior region showing slit; (D-F) G. obtura paratype MZSP 121837 (L 36.4 mm), same views as preceding specimen; (G-I) G. semistriolata MZSP 47357 (L 26.4 mm),same views as first specimen.Scales:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14705714" httpUri="https://zenodo.org/record/14705714/files/figure.png" pageId="96" pageNumber="97">Fig. 64C, F</figureCitation>
), from 11 to 15% of shell length, sometimes longitudinal (
<figureCitation id="509A2A1DFFA9A8448CBDDD16FB41FD81" box="[1160,1255,570,595]" captionStart="Figure 64" captionStartId="96.[170,229,2042,2067]" captionTargetBox="[170,1493,859,2012]" captionTargetId="figure-228@96.[170,1493,859,2012]" captionTargetPageId="96" captionText="Figure 64. Shell of types of Graptacme obtura and G. semistriolata: (A) G. obtura holotype MZSP 166760 (L 35.6 mm), dorsal view; (B) same, right view; (C) same, detail of posterior region showing slit; (D-F) G. obtura paratype MZSP 121837 (L 36.4 mm), same views as preceding specimen; (G-I) G. semistriolata MZSP 47357 (L 26.4 mm),same views as first specimen.Scales:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14705714" httpUri="https://zenodo.org/record/14705714/files/figure.png" pageId="96" pageNumber="97">Fig. 64F</figureCitation>
), sometimes slightly oblique (
<figureCitation id="509A2A1DFFA9A8448BFCDD76FB81FDA1" box="[969,1063,602,627]" captionStart="Figure 64" captionStartId="96.[170,229,2042,2067]" captionTargetBox="[170,1493,859,2012]" captionTargetId="figure-228@96.[170,1493,859,2012]" captionTargetPageId="96" captionText="Figure 64. Shell of types of Graptacme obtura and G. semistriolata: (A) G. obtura holotype MZSP 166760 (L 35.6 mm), dorsal view; (B) same, right view; (C) same, detail of posterior region showing slit; (D-F) G. obtura paratype MZSP 121837 (L 36.4 mm), same views as preceding specimen; (G-I) G. semistriolata MZSP 47357 (L 26.4 mm),same views as first specimen.Scales:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14705714" httpUri="https://zenodo.org/record/14705714/files/figure.png" pageId="96" pageNumber="97">Fig. 64C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="80BB6513FFA9A8448B50DDB6FC10FD21" pageId="96" pageNumber="97" type="etymology">
<paragraph id="C81E3698FFA9A8448B50DDB6FC10FD21" blockId="96.[869,1493,666,755]" pageId="96" pageNumber="97">
<emphasis id="FAD5EA8AFFA9A8448B50DDB6FC4BFD61" bold="true" box="[869,1005,666,691]" pageId="96" pageNumber="97">Etymology:</emphasis>
The specific epithet is derived from the Latin word 
<emphasis id="FAD5EA8AFFA9A8448BF7DD96FBBCFD01" box="[962,1050,698,723]" italics="true" pageId="96" pageNumber="97">obturus,</emphasis>
meaning &quot;plug&quot;, an allusion to the shell format.
</paragraph>
</subSubSection>
<subSubSection id="80BB6513FFA9A8448B50DC36FA3FFCE1" box="[869,1433,794,819]" pageId="96" pageNumber="97" type="distribution">
<paragraph id="C81E3698FFA9A8448B50DC36FA3FFCE1" blockId="96.[869,1433,794,819]" box="[869,1433,794,819]" pageId="96" pageNumber="97">
<emphasis id="FAD5EA8AFFA9A8448B50DC36FC5BFCE1" bold="true" box="[869,1021,794,819]" pageId="96" pageNumber="97">Distribution:</emphasis>
Endemic from Trindade Island, 
<collectingCountry id="B0B67608FFA9A8448D62DC36FA33FCE1" box="[1367,1429,794,819]" name="Brazil" pageId="96" pageNumber="97">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>