<document id="466D187D855B0D859DA08DE797F3F97D" ID-DOI="10.11606/1807-0205/2024.64.031" ID-ISSN="1807-0205" ID-Zenodo-Dep="14705562" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737339390010" checkinUser="felipe" docAuthor="Simone, Luiz Ricardo L." docDate="2024" docId="4008878EFFC2A82888F1DBB6FDCEFE81" docLanguage="en" docName="PapAvZool.64.e202464031.pdf" docOrigin="Papéis Avulsos de Zoologia 64" docSource="https://doi.org/10.11606/1807-0205/2024.64.031" docStyle="DocumentStyle:6844D5473582683DD13A9A75B001759F.4:PapAvZool.2018-.journal_article" docStyleId="6844D5473582683DD13A9A75B001759F" docStyleName="PapAvZool.2018-.journal_article" docStyleVersion="4" docTitle="Depressiscala nautlae" docType="treatment" docVersion="2" lastPageNumber="12" masterDocId="BC31FFF6FFC9A8248835DF2CFFA6FFD2" masterDocTitle="New species, misidentifications and problematic taxonomy of some Atlantic South American marine mollusks: a review" masterLastPageNumber="104" masterPageNumber="1" pageNumber="12" updateTime="1737393675899" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0" zenodo-license-figures="CC-BY-NC-4.0">
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<paragraph id="C81E3698FFC2A82F88F1DBB6FE74FB01" blockId="11.[196,617,986,1235]" pageId="11" pageNumber="12">
<emphasis id="FAD5EA8AFFC2A82F88F1DBB6FE74FB01" bold="true" pageId="11" pageNumber="12">
<taxonomicName id="0FA14D1BFFC2A82F88F1DBB6FDCFFB61" ID-CoL="34V49" authority="(Morch, 1875)" baseAuthorityName="Morch" baseAuthorityYear="1875" box="[196,617,1178,1203]" class="Gastropoda" family="Epitoniidae" genus="Depressiscala" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="nautlae">
<emphasis id="FAD5EA8AFFC2A82F88F1DBB6FE66FB61" bold="true" box="[196,448,1178,1203]" italics="true" pageId="11" pageNumber="12">Depressiscala nautlae</emphasis>
(
<bibRefCitation id="AC304B69FFC2A82F89F9DBB6FDC7FB61" author="Morch, O. A. L." box="[460,609,1178,1203]" pageId="11" pageNumber="12" pagination="250 - 268" refId="ref79997" refString="Morch, O. A. L. 1875. Synopsis familiae Scalidarum Indiarum occidentalium. Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjobenhavn, 1874 (17): 250 - 268." type="journal article" year="1875">Mörch, 1875</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="509A2A1DFFC2A82F8957DB96FE06FB01" box="[354,416,1210,1235]" captionStart="Figure 7" captionStartId="11.[94,153,2042,2067]" captionTargetBox="[94,1417,1351,2012]" captionTargetId="figure-569@11.[94,1417,1351,2012]" captionTargetPageId="11" captionText="Figure 7. Depressiscala nautlae and D. nitidella shells: (A-B) D. nautlae, MZSP 166500, from Ubatuba, São Paulo, Brazil (L 14.0 mm), frontal and dorsal views; (C) same, MZSP 165694, same locality (L 13.1 mm); (D-E) Lectotype of Scala scipio USNM 10694 (L 14.9 mm); (F) D. nautlae (L 13.3 mm), ordinary specimen from Pompano Beach,Florida,ANSP 457201, frontal view;(G) Scala nitidella, lectotypeUSNM 83716 (L 13.1 mm), frontal view (USNM photos courtesy Ellen Strong)." figureDoi="http://doi.org/10.5281/zenodo.14705581" httpUri="https://zenodo.org/record/14705581/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
A-F)
</emphasis>
</paragraph>
<paragraph id="C81E3698FFC2A82F886BDBD6FE1FFAC1" blockId="11.[94,441,1274,1331]" box="[94,441,1274,1299]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="E8B111B6FFC2A82F886BDBD6FE1FFAC1" box="[94,441,1274,1299]" pageId="11" pageNumber="12">
<treatmentCitation id="49001089FFC2A82F886BDBD6FE10FAC1" author="Morch, O. A. L." box="[94,438,1274,1299]" page="265" pageId="11" pageNumber="12" year="1875">
<taxonomicName id="0FA14D1BFFC2A82F886BDBD6FE10FAC1" authority="Morch, 1875: 265" authorityName="Morch" authorityPageNumber="265" authorityYear="1875" box="[94,438,1274,1299]" class="Gastropoda" family="Vanikoridae" genus="Scala" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="nautlae">
<emphasis id="FAD5EA8AFFC2A82F886BDBD6FF56FAC1" box="[94,240,1274,1299]" italics="true" pageId="11" pageNumber="12">Scala nautlae</emphasis>
<bibRefCitation id="AC304B69FFC2A82F88C0DBD6FE10FAC1" author="Morch, O. A. L." box="[245,438,1274,1299]" pageId="11" pageNumber="12" pagination="250 - 268" refId="ref79997" refString="Morch, O. A. L. 1875. Synopsis familiae Scalidarum Indiarum occidentalium. Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjobenhavn, 1874 (17): 250 - 268." type="journal article" year="1875">Mörch, 1875: 265</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C81E3698FFC2A82F886BDA36FACBFEE1" blockId="11.[94,441,1274,1331]" lastBlockId="11.[794,1419,218,339]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="E8B111B6FFC2A82F886BDA36FACBFEE1" pageId="11" pageNumber="12">
<treatmentCitation id="49001089FFC2A82F886BDA36FED0FAE1" author="Dall, W. H." box="[94,374,1306,1331]" page="27" pageId="11" pageNumber="12" year="1889">
<taxonomicName id="0FA14D1BFFC2A82F886BDA36FED0FAE1" authority="Dall, 1889: 27" authorityName="Dall" authorityPageNumber="27" authorityYear="1889" box="[94,374,1306,1331]" class="Gastropoda" family="Vanikoridae" genus="Scala" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="scipio">
<emphasis id="FAD5EA8AFFC2A82F886BDA36FF7CFAE1" box="[94,218,1306,1331]" italics="true" pageId="11" pageNumber="12">Scala scipio</emphasis>
<bibRefCitation id="AC304B69FFC2A82F88D5DA36FED0FAE1" author="Dall, W. H." box="[224,374,1306,1331]" pageId="11" pageNumber="12" pagination="1 - 492" refId="ref77257" refString="Dall, W. H. 1889. Reports on the results of dredgings, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer ' Blake'. Bulletin of the Museum of Comparative Zoology, 18: 1 - 492, pls. 10 - 40." type="journal article" year="1889">Dall, 1889: 27</bibRefCitation>
</taxonomicName>
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, 310. 
<emphasis id="FAD5EA8AFFC2A82F8B2FDFF6FB33FF21" box="[794,1173,218,243]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="0FA14D1BFFC2A82F8B2FDFF6FB36FF21" ID-CoL="6G9CP" baseAuthorityName="Morch" baseAuthorityYear="1875" box="[794,1168,218,243]" class="Gastropoda" family="Epitoniidae" genus="Epitonium" kingdom="Animalia" order="Architaenioglossa" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="nautlae" subGenus="Depressiscala">Epitonium (Depressiscala) nautlae</taxonomicName>
:
</emphasis>
<treatmentCitation id="49001089FFC2A82F8C9EDFF6FA2FFF21" author="Abbott, R. T." box="[1195,1417,218,243]" page="122" pageId="11" pageNumber="12" year="1974">
<bibRefCitation id="AC304B69FFC2A82F8C9EDFF6FA2FFF21" author="Abbott, R. T." box="[1195,1417,218,243]" pageId="11" pageNumber="12" refId="ref75583" refString="Abbott, R. T. 1974. American Seashells, second edition. Van Nostrand Reinhold Company. New York, 663 pp., 240 pls." type="book" year="1974">Abbott, 1974: 122</bibRefCitation>
</treatmentCitation>
(fig.1271); 
<treatmentCitation id="49001089FFC2A82F8B9BDFD6FB9CFEC1" author="Rios, E. C." box="[942,1082,250,275]" page="56" pageId="11" pageNumber="12" year="1975">
<bibRefCitation id="AC304B69FFC2A82F8B9BDFD6FB9CFEC1" author="Rios, E. C." box="[942,1082,250,275]" pageId="11" pageNumber="12" refId="ref81445" refString="Rios, E. C. 1975. Brazilian marine mollusks iconography. Fundacao Cidade do Rio Grande. Rio Grande, 331 pp., 91 pls." type="book" year="1975">Rios,1975:56</bibRefCitation>
</treatmentCitation>
(pl.15,fig.217),1994:157(pl.52, fig.743), 1994:99 (pl. 32,fig. 400),2009:181 (fig.439).
</treatmentCitationGroup>
</paragraph>
<paragraph id="C81E3698FFC2A82F8B2FDE16FA9CFE81" blockId="11.[794,1419,218,339]" box="[794,1338,314,339]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="E8B111B6FFC2A82F8B2FDE16FA9CFE81" box="[794,1338,314,339]" pageId="11" pageNumber="12">
<emphasis id="FAD5EA8AFFC2A82F8B2FDE16FBAEFE81" box="[794,1032,314,339]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="0FA14D1BFFC2A82F8B2FDE16FBA4FE81" ID-CoL="34V49" baseAuthorityName="Morch" baseAuthorityYear="1875" box="[794,1026,314,339]" class="Gastropoda" family="Epitoniidae" genus="Depressiscala" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="nautlae">Depressiscala nautlae</taxonomicName>
:
</emphasis>
<treatmentCitation id="49001089FFC2A82F8C38DE16FA93FE81" author="Rosenberg, G. &amp; Moretzsohn, F. &amp; Garcia, E. F." box="[1037,1333,314,339]" page="641" pageId="11" pageNumber="12" year="2009">
<bibRefCitation id="AC304B69FFC2A82F8C38DE16FA93FE81" author="Rosenberg, G. &amp; Moretzsohn, F. &amp; Garcia, E. F." box="[1037,1333,314,339]" pageId="11" pageNumber="12" pagination="579 - 699" refId="ref81599" refString="Rosenberg, G.; Moretzsohn, F. &amp; Garcia, E. F. 2009. Gastropoda (Mollusca) of the Gulf of Mexico, pp. 579 - 699. In: Felder, D. L. &amp; Camp, D. K. (Eds.). Gulf of Mexicoorigins, waters, and biota. Biodiversity. Texas. A &amp; M Press. College Station. Texas." type="book chapter" year="2009">
Rosenberg 
<emphasis id="FAD5EA8AFFC2A82F8CBEDE16FB65FE81" box="[1163,1219,314,339]" italics="true" pageId="11" pageNumber="12">et al.,</emphasis>
2009: 641
</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="80BB6513FFC2A82F8B2FDE56FABAFE01" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="C81E3698FFC2A82F8B2FDE56FABAFE01" blockId="11.[794,1418,378,467]" pageId="11" pageNumber="12">
<emphasis id="FAD5EA8AFFC2A82F8B2FDE56FC6FFE41" bold="true" box="[794,969,378,403]" pageId="11" pageNumber="12">
<typeStatus id="171A883AFFC2A82F8B2FDE56FCF5FE41" box="[794,851,378,403]" pageId="11" pageNumber="12">Type</typeStatus>
localities:
</emphasis>
<emphasis id="FAD5EA8AFFC2A82F8BFADE56FB98FE41" box="[975,1086,378,403]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="0FA14D1BFFC2A82F8BFADE56FB9EFE41" baseAuthorityName="Morch" baseAuthorityYear="1875" box="[975,1080,378,403]" class="Gastropoda" family="Epitoniidae" genus="Depressiscala" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="nautlae">S.nautlae</taxonomicName>
:
</emphasis>
ad litus mexicanum inter Vera Cruz et Nautla. 
<emphasis id="FAD5EA8AFFC2A82F8BF6DEB6FBE3FE61" box="[963,1093,410,435]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="0FA14D1BFFC2A82F8BF6DEB6FBE6FE61" authorityName="Dall" authorityYear="1889" box="[963,1088,410,435]" class="Gastropoda" family="Vanikoridae" genus="Scala" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="scipio">Scala scipio</taxonomicName>
:
</emphasis>
Vera Cruz; sta. 2597, 
<quantity id="0F599B7DFFC2A82F8D18DEB6FA2CFE61" box="[1325,1418,410,435]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="11" pageNumber="12" unit="mi" value="20.0">20 miles</quantity>
SW by S from 
<collectingRegion id="0A65F87AFFC2A82F8B87DE96FC4DFE01" box="[946,1003,442,467]" country="South Africa" name="Western Cape" pageId="11" pageNumber="12">Cape</collectingRegion>
Hattenas, N.C., 15 + 12 fms.
</paragraph>
</subSubSection>
<subSubSection id="80BB6513FFC2A82F8B2FDED6FB33FB01" pageId="11" pageNumber="12" type="discussion">
<paragraph id="C81E3698FFC2A82F8B2FDED6FB36FCC1" blockId="11.[794,1419,506,1235]" pageId="11" pageNumber="12">
<emphasis id="FAD5EA8AFFC2A82F8B2FDED6FC2EFDC1" bold="true" box="[794,904,506,531]" pageId="11" pageNumber="12">Remarks:</emphasis>
Specimens found on the Brazilian coast with the shell characteristics shown in 
<figureCitation id="509A2A1DFFC2A82F8C95DD36FB7AFDE1" box="[1184,1244,538,563]" captionStart="Figure 7" captionStartId="11.[94,153,2042,2067]" captionTargetBox="[94,1417,1351,2012]" captionTargetId="figure-569@11.[94,1417,1351,2012]" captionTargetPageId="11" captionText="Figure 7. Depressiscala nautlae and D. nitidella shells: (A-B) D. nautlae, MZSP 166500, from Ubatuba, São Paulo, Brazil (L 14.0 mm), frontal and dorsal views; (C) same, MZSP 165694, same locality (L 13.1 mm); (D-E) Lectotype of Scala scipio USNM 10694 (L 14.9 mm); (F) D. nautlae (L 13.3 mm), ordinary specimen from Pompano Beach,Florida,ANSP 457201, frontal view;(G) Scala nitidella, lectotypeUSNM 83716 (L 13.1 mm), frontal view (USNM photos courtesy Ellen Strong)." figureDoi="http://doi.org/10.5281/zenodo.14705581" httpUri="https://zenodo.org/record/14705581/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
A-C were identified as both 
<taxonomicName id="0FA14D1BFFC2A82F8B88DD16FB88FD81" baseAuthorityName="Morch" baseAuthorityYear="1875" box="[957,1070,570,595]" class="Gastropoda" family="Epitoniidae" genus="Depressiscala" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="nautlae">
<emphasis id="FAD5EA8AFFC2A82F8B88DD16FB88FD81" box="[957,1070,570,595]" italics="true" pageId="11" pageNumber="12">D. nautlae</emphasis>
</taxonomicName>
and 
<taxonomicName id="0FA14D1BFFC2A82F8C5EDD16FACAFD81" authority="(Dall, 1889)" baseAuthorityName="Dall" baseAuthorityYear="1889" box="[1131,1388,570,595]" class="Gastropoda" family="Epitoniidae" genus="Depressiscala" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="niditella">
<emphasis id="FAD5EA8AFFC2A82F8C5EDD16FB47FD81" box="[1131,1249,570,595]" italics="true" pageId="11" pageNumber="12">D. niditella</emphasis>
(
<bibRefCitation id="AC304B69FFC2A82F8CC4DD16FAC1FD81" author="Dall, W. H." box="[1265,1383,570,595]" pageId="11" pageNumber="12" pagination="1 - 492" refId="ref77257" refString="Dall, W. H. 1889. Reports on the results of dredgings, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877 - 78) and in the Caribbean Sea (1879 - 80), by the U. S. Coast Survey Steamer ' Blake'. Bulletin of the Museum of Comparative Zoology, 18: 1 - 492, pls. 10 - 40." type="journal article" year="1889">Dall, 1889</bibRefCitation>
)
</taxonomicName>
. A comparison to the 
<typeStatus id="171A883AFFC2A82F8BDFDD76FBF5FDA1" box="[1002,1107,602,627]" pageId="11" pageNumber="12" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="0FA14D1BFFC2A82F8C41DD76FB40FDA1" baseAuthorityName="Dall" baseAuthorityYear="1889" box="[1140,1254,602,627]" class="Gastropoda" family="Epitoniidae" genus="Depressiscala" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="nitidella">
<emphasis id="FAD5EA8AFFC2A82F8C41DD76FB40FDA1" box="[1140,1254,602,627]" italics="true" pageId="11" pageNumber="12">D. nitidella</emphasis>
</taxonomicName>
(USNM 83716) (
<figureCitation id="509A2A1DFFC2A82F8B15DD56FCC9FD41" box="[800,879,634,659]" captionStart="Figure 7" captionStartId="11.[94,153,2042,2067]" captionTargetBox="[94,1417,1351,2012]" captionTargetId="figure-569@11.[94,1417,1351,2012]" captionTargetPageId="11" captionText="Figure 7. Depressiscala nautlae and D. nitidella shells: (A-B) D. nautlae, MZSP 166500, from Ubatuba, São Paulo, Brazil (L 14.0 mm), frontal and dorsal views; (C) same, MZSP 165694, same locality (L 13.1 mm); (D-E) Lectotype of Scala scipio USNM 10694 (L 14.9 mm); (F) D. nautlae (L 13.3 mm), ordinary specimen from Pompano Beach,Florida,ANSP 457201, frontal view;(G) Scala nitidella, lectotypeUSNM 83716 (L 13.1 mm), frontal view (USNM photos courtesy Ellen Strong)." figureDoi="http://doi.org/10.5281/zenodo.14705581" httpUri="https://zenodo.org/record/14705581/files/figure.png" pageId="11" pageNumber="12">Fig. 7G</figureCitation>
) reveals that they do not belong to that species, as the shells are much slenderer and lack an umbilicus. Conversely, their shells are virtually identical to the 
<typeStatus id="171A883AFFC2A82F8D7FDD96FCEAFD21" pageId="11" pageNumber="12" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="0FA14D1BFFC2A82F8B5EDDF6FC40FD21" authorityName="Dall" authorityYear="1889" box="[875,998,730,755]" class="Gastropoda" family="Vanikoridae" genus="Scala" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="scipio">
<emphasis id="FAD5EA8AFFC2A82F8B5EDDF6FC40FD21" box="[875,998,730,755]" italics="true" pageId="11" pageNumber="12">Scala scipio</emphasis>
</taxonomicName>
(
<figureCitation id="509A2A1DFFC2A82F8BC4DDF6FBF0FD21" box="[1009,1110,730,755]" captionStart="Figure 7" captionStartId="11.[94,153,2042,2067]" captionTargetBox="[94,1417,1351,2012]" captionTargetId="figure-569@11.[94,1417,1351,2012]" captionTargetPageId="11" captionText="Figure 7. Depressiscala nautlae and D. nitidella shells: (A-B) D. nautlae, MZSP 166500, from Ubatuba, São Paulo, Brazil (L 14.0 mm), frontal and dorsal views; (C) same, MZSP 165694, same locality (L 13.1 mm); (D-E) Lectotype of Scala scipio USNM 10694 (L 14.9 mm); (F) D. nautlae (L 13.3 mm), ordinary specimen from Pompano Beach,Florida,ANSP 457201, frontal view;(G) Scala nitidella, lectotypeUSNM 83716 (L 13.1 mm), frontal view (USNM photos courtesy Ellen Strong)." figureDoi="http://doi.org/10.5281/zenodo.14705581" httpUri="https://zenodo.org/record/14705581/files/figure.png" pageId="11" pageNumber="12">Fig. 7D, E</figureCitation>
), a species presently considered synonymous with 
<emphasis id="FAD5EA8AFFC2A82F8C29DDD6FB36FCC1" box="[1052,1168,762,787]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="0FA14D1BFFC2A82F8C29DDD6FB2AFCC1" baseAuthorityName="Morch" baseAuthorityYear="1875" box="[1052,1164,762,787]" class="Gastropoda" family="Epitoniidae" genus="Depressiscala" kingdom="Animalia" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="nautlae">D. nautlae</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="C81E3698FFC2A82F8B0ADC36FB33FB01" blockId="11.[794,1419,506,1235]" pageId="11" pageNumber="12">
The issue arises from the apparent existence of two different morphs under this epithet. One resembles the 
<taxonomicName id="0FA14D1BFFC2A82F8B2FDC76FCD6FCA1" authorityName="Dall" authorityYear="1889" box="[794,880,858,883]" class="Gastropoda" family="Vanikoridae" genus="Scala" kingdom="Animalia" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="scipio">
<emphasis id="FAD5EA8AFFC2A82F8B2FDC76FCD6FCA1" box="[794,880,858,883]" italics="true" pageId="11" pageNumber="12">S. scipio</emphasis>
</taxonomicName>
model, with a deeper suture, more distant and narrower axial threads, and always with an amputated apex (
<figureCitation id="509A2A1DFFC2A82F8B6BDCB6FC3AFC61" box="[862,924,922,947]" captionStart="Figure 7" captionStartId="11.[94,153,2042,2067]" captionTargetBox="[94,1417,1351,2012]" captionTargetId="figure-569@11.[94,1417,1351,2012]" captionTargetPageId="11" captionText="Figure 7. Depressiscala nautlae and D. nitidella shells: (A-B) D. nautlae, MZSP 166500, from Ubatuba, São Paulo, Brazil (L 14.0 mm), frontal and dorsal views; (C) same, MZSP 165694, same locality (L 13.1 mm); (D-E) Lectotype of Scala scipio USNM 10694 (L 14.9 mm); (F) D. nautlae (L 13.3 mm), ordinary specimen from Pompano Beach,Florida,ANSP 457201, frontal view;(G) Scala nitidella, lectotypeUSNM 83716 (L 13.1 mm), frontal view (USNM photos courtesy Ellen Strong)." figureDoi="http://doi.org/10.5281/zenodo.14705581" httpUri="https://zenodo.org/record/14705581/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
A-E). The other morph, illustrated in 
<figureCitation id="509A2A1DFFC2A82F8D09DCB6FA21FC61" box="[1340,1415,922,947]" captionStart="Figure 7" captionStartId="11.[94,153,2042,2067]" captionTargetBox="[94,1417,1351,2012]" captionTargetId="figure-569@11.[94,1417,1351,2012]" captionTargetPageId="11" captionText="Figure 7. Depressiscala nautlae and D. nitidella shells: (A-B) D. nautlae, MZSP 166500, from Ubatuba, São Paulo, Brazil (L 14.0 mm), frontal and dorsal views; (C) same, MZSP 165694, same locality (L 13.1 mm); (D-E) Lectotype of Scala scipio USNM 10694 (L 14.9 mm); (F) D. nautlae (L 13.3 mm), ordinary specimen from Pompano Beach,Florida,ANSP 457201, frontal view;(G) Scala nitidella, lectotypeUSNM 83716 (L 13.1 mm), frontal view (USNM photos courtesy Ellen Strong)." figureDoi="http://doi.org/10.5281/zenodo.14705581" httpUri="https://zenodo.org/record/14705581/files/figure.png" pageId="11" pageNumber="12">Fig. 7F</figureCitation>
, features a shallower suture, taller and more closely locat- ed axial threads, and a complete apex. This latter morph is illustrated, for example, in 
<bibRefCitation id="AC304B69FFC2A82F8C50DCD6FAA3FBC1" author="Abbott, R. T." box="[1125,1285,1018,1043]" pageId="11" pageNumber="12" refId="ref75583" refString="Abbott, R. T. 1974. American Seashells, second edition. Van Nostrand Reinhold Company. New York, 663 pp., 240 pls." type="book" year="1974">Abbott (1974)</bibRefCitation>
. Upon analyzing these specimens, the most parsimonious conclusion is that they belong to different species. However, it is important to determine whether the 
<typeStatus id="171A883AFFC2A82F8CE8DB76FABCFBA1" box="[1245,1306,1114,1139]" pageId="11" pageNumber="12">types</typeStatus>
of 
<bibRefCitation id="AC304B69FFC2A82F8D77DB76FCF9FB41" author="Morch, O. A. L." pageId="11" pageNumber="12" pagination="250 - 268" refId="ref79997" refString="Morch, O. A. L. 1875. Synopsis familiae Scalidarum Indiarum occidentalium. Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjobenhavn, 1874 (17): 250 - 268." type="journal article" year="1875">Mörch (1875)</bibRefCitation>
resemble them, as they have not been located or photographed to the best of our knowledge. This is an important issue for future surveys.
</paragraph>
</subSubSection>
<subSubSection id="80BB6513FFC2A8288B2FDBD6FDCEFE81" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="C81E3698FFC2A8288B2FDBD6FDCEFE81" blockId="11.[794,1419,1274,1331]" lastBlockId="12.[170,795,218,339]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="FAD5EA8AFFC2A82F8B2FDBD6FBAEFAC1" bold="true" box="[794,1032,1274,1299]" pageId="11" pageNumber="12">Material examined:</emphasis>
<materialsCitation id="78C93CC5FFC2A82F8C2DDBD6FB16FAC1" box="[1048,1200,1274,1299]" collectorName="Florida" country="United States of America" location="USA" pageId="11" pageNumber="12" specimenCount="1">
<collectingCountry id="B0B67608FFC2A82F8C2DDBD6FBECFAC1" box="[1048,1098,1274,1299]" name="United States of America" pageId="11" pageNumber="12">USA</collectingCountry>
. 
<emphasis id="FAD5EA8AFFC2A82F8C69DBD6FB16FAC1" bold="true" box="[1116,1200,1274,1299]" pageId="11" pageNumber="12">
<collectorName id="6554534EFFC2A82F8C69DBD6FB16FAC1" box="[1116,1200,1274,1299]" pageId="11" pageNumber="12">Florida</collectorName>
</emphasis>
</materialsCitation>
; 
<materialsCitation id="78C93CC5FFC2A8288CF0DBD6FEFDFF21" collectionCode="ANSP" collectorName="Pompano Beach &amp; Harriet &amp; Robertson &amp; Golberg" country="United States of America" elevation="52" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" specimenCode="ANSP 457201, 2" specimenCount="1">
<collectorName id="6554534EFFC2A82F8CF0DBD6FA20FAC1" box="[1221,1414,1274,1299]" pageId="11" pageNumber="12">Pompano Beach</collectorName>
, 
<quantity id="0F599B7DFFC2A82F8B2FDA36FCF0FAE1" box="[794,854,1306,1331]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="11" pageNumber="12" unit="m" value="52.0">
<elevation id="438CD1ABFFC2A82F8B2FDA36FCF0FAE1" box="[794,854,1306,1331]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="11" pageNumber="12" unit="m" value="52.0">52 m</elevation>
</quantity>
, 
<specimenCode id="98079EE3FFC2A82F8B57DA36FC5AFAE1" box="[866,1020,1306,1331]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="11" pageNumber="12">ANSP 457201</specimenCode>
, 
<specimenCode id="98079EE3FFC2A82F8C3DDA36FBB0FAE1" box="[1032,1046,1306,1331]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="11" pageNumber="12">2</specimenCode>
shells (
<collectorName id="6554534EFFC2A82F8C5FDA36FB1CFAE1" box="[1130,1210,1306,1331]" pageId="11" pageNumber="12">Harriet</collectorName>
, 
<collectorName id="6554534EFFC2A82F8CF3DA36FA9CFAE1" box="[1222,1338,1306,1331]" pageId="11" pageNumber="12">Robertson</collectorName>
&amp; 
<collectorName id="6554534EFFC2A8288D6EDA36FF78FF21" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">Golberg</collectorName>
col., 2004)
</materialsCitation>
. 
<materialsCitation id="78C93CC5FFC5A8288953DFF6FD9FFF21" box="[358,569,218,243]" country="Brazil" location="Sao Paulo" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Sao Paulo">
<collectingCountry id="B0B67608FFC5A8288953DFF6FE1EFF21" box="[358,440,218,243]" name="Brazil" pageId="12" pageNumber="13">BRAZIL</collectingCountry>
. 
<emphasis id="FAD5EA8AFFC5A82889F7DFF6FD9FFF21" bold="true" box="[450,569,218,243]" pageId="12" pageNumber="13">
<collectingRegion id="0A65F87AFFC5A82889F7DFF6FD9FFF21" box="[450,569,218,243]" country="Brazil" name="Sao Paulo" pageId="12" pageNumber="13">São Paulo</collectingRegion>
</emphasis>
</materialsCitation>
; 
<materialsCitation id="78C93CC5FFC5A8288A72DFF6FDC2FE81" collectingDate="2023-08" collectionCode="MZSP" collectorName="Ubatuba &amp; D. Forcelli &amp; Plus" country="Brazil" latitude="-23.352556" location="Piaui" longLatPrecision="1" longitude="-44.93175" pageId="12" pageNumber="13" specimenCode="MZSP 166500, 1" specimenCount="1" stateProvince="Piaui">
<collectorName id="6554534EFFC5A8288A72DFF6FD0CFF21" box="[583,682,218,243]" pageId="12" pageNumber="13">Ubatuba</collectorName>
, 
<taxonomicName id="0FA14D1BFFC5A8288A80DFF6FF46FEC1" authority="Puruba" authorityName="Puruba" class="Bivalvia" family="Ostreidae" genus="Crassostrea" kingdom="Animalia" order="Ostreida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="praia">praia Puruba</taxonomicName>
, 
<geoCoordinate id="AD95505FFFC5A82888D9DFD6FED1FEC1" box="[236,375,249,275]" degrees="23" direction="south" minutes="21" orientation="latitude" pageId="12" pageNumber="13" precision="1" seconds="09.2" value="-23.352556">23°21′09.2″S</geoCoordinate>
<geoCoordinate id="AD95505FFFC5A82889B5DFD6FDB0FEC1" box="[384,534,249,275]" degrees="44" direction="west" minutes="55" orientation="longitude" pageId="12" pageNumber="13" precision="1" seconds="54.3" value="-44.93175">44°55′54.3″W</geoCoordinate>
, 
<specimenCode id="98079EE3FFC5A8288A17DFD6FD19FEC1" box="[546,703,250,275]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13" type="Museum">MZSP 166500</specimenCode>
, 
<specimenCode id="98079EE3FFC5A8288AFEDFD6FD7FFEC1" box="[715,729,250,275]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13" type="Museum">1</specimenCode>
shell, 165694, 7 shells (
<collectorName id="6554534EFFC5A828895CDE36FE73FEE1" box="[361,469,282,307]" pageId="12" pageNumber="13">D. Forcelli</collectorName>
col., 
<date id="BC1F1058FFC5A8288A3BDE36FDC8FEE1" box="[526,622,282,307]" pageId="12" pageNumber="13" value="2023-08">
<collectingDate id="AC5BE9B0FFC5A8288A3BDE36FDC8FEE1" box="[526,622,282,307]" pageId="12" pageNumber="13" value="2023-08">viii.2023</collectingDate>
</date>
). 
<collectorName id="6554534EFFC5A8288A4BDE36FD0AFEE1" box="[638,684,282,307]" pageId="12" pageNumber="13">Plus</collectorName>
11 lots of 
<collectionCode id="AEB0AE5DFFC5A828889FDE16FF4CFE81" box="[170,234,314,339]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="12" pageNumber="13" type="Museum">MZSP</collectionCode>
collection from 
<collectingRegion id="0A65F87AFFC5A82889A8DE16FE75FE81" box="[413,467,314,339]" country="Brazil" name="Piaui" pageId="12" pageNumber="13">Piauí</collectingRegion>
to 
<collectingRegion id="0A65F87AFFC5A82889C0DE16FDC2FE81" box="[501,612,314,339]" country="Brazil" name="Sao Paulo" pageId="12" pageNumber="13">São Paulo</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>