There and back again: More on the taxonomy of the crab spiders genus Epicadus (Thomisidae: Stephanopinae) Author Machado, Miguel Author Teixeira, Renato Augusto Author Lise, Arno Antonio text Zootaxa 2018 2018-02-22 4382 3 501 530 journal article 30682 10.11646/zootaxa.4382.3.4 b88bfa2d-4b83-41f8-a145-a95ca196960f 1175-5326 1183130 87A2B333-09FF-4726-A1EF-0699FB631808 Epicadus pustulosus ( Mello-Leitão, 1929 ) Figs 14A–D , 15E Tobias pustulosus Mello-Leitão, 1929 : 86 , figs 183–184. Machado et al. 2015 : 572 , figs 12–24. Tobias monstrosus Simon in Mello-Leitão, 1929 : 87 , figs 185–186. Epicadus pustulosus (Mello-Leitão) . Machado et al. 2017 : 448 , figs S7D, S 8F. Type material: Holotype (original designation): subadult female, Fonte Boa [ 2°28'5.16"S , 66°8'29.49"W , Amazonas , Brazil ] ( MNHN 4560 , examined). Other material examined. BRAZIL : Acre : 1 male , Senador Guiomard ( Reserva Extrativista de Catuaba ), 10°08'43"S , 67°43'29"W , 2002, E.F. Morato ( IBSP 135286 ) ; 1 male (IBSP 135287); 1 male (IBSP 135289); 1 male (IBSP 135290); 1 female (IBSP 135291); 1 male ( IBSP 135292 ); all same locality and collection data . Rondônia : 1 female , Porto Velho , 8°48'10"S , 63°55'05"W , 20 January 2012 , D.F. Candiani ( MZSP 45458 ) ; 1 female, 4 July 2012 , A.A. Nogueira (MZSP 51068); 1 female , 8 July 2012 , R.P. Indicatti ( MZSP 53085 .; Bahia : 1 female , Gandu, Fazenda São Rafael (cocoa forest), 13°44'38"S , 39°29'05"W , 5 November 1968 ( MNRJ 1351 ) ; 1 male , Una , Reserva Biológica de Una , 15°12'32"S , 39°11'45"W , 15–28 November 2000 , A.D. Brescovit et al. ( IBSP 46728 ) . Espírito Santo : 1 male , São Mateus , Reserva Floresta da Cia. Vale do Rio Doce , 19°09'05"S , 40°04'14"W , July 1997 , A.J. Santos ( IBSP 27015 ) . São Paulo : 1 female , São Paulo , Parque Modelo , 22°42'24"S , 46°48'34"W , August 1958 , M. Kramer ( IBSP 137989 ) ; 1 male , Mogi das Cruzes , Manuel Ferreira , Birituba-Açu , 23°29'22"S , 46°09'59"W , May 2001 , E.K. Kashimata & R. Martins ( IBSP 56568 ) ; 1 female , Parque Estadual da Ilha do Cardoso , 25°8'1.02"S , 47°57'41.94"W , 23–25 August 1999 , G. Machado ( MZSP 21592 ) . ECUADOR : Cotopaxi : 1 male , Rio Napo , 0°59'46"S , 78°25'18"W , January 1972 , Bordon ( MACN 18958 ). Diagnosis. Females of E. pustulosus are very similar to those of E. caudatus in their cryptic bark-dweller habitus with rugose tegument and predominant dark-brown coloration. Both species also share squared leg segments and protuberant anterior femora suffused with many conical tubercles. However, E. pustulosus can be distinguished by the yellow blister-shaped sockets on the ventral surface of femora I and II and by the anterior projection on the epigynal plate, which covers the copulatory openings entirely. Males can be easily recognized by their equal-sized, short and spiniform opisthosomal projections, by a basal indentation on the DTA and by its curvature (pointing to the dorsum of the cymbium). Description. See Machado et al. (2015) . Distribution. BRAZIL : São Paulo , Espiírito Santo, Bahia ; ECUADOR : Cotopaxi ; PANAMA : Panama City , Colón ( Fig. 17A ). Variation. Most of the specimens examined by Machado et al. (2015) present a predominantly brown coloration. Here, we present two additional color variations for the females of E. pustulosus : dorsum of prosoma and opisthosoma black with front and legs yellow ( Figs 14A, B ) and brown with distal portion of femora I and dorsum of opisthosoma yellow ( Figs 14C, D ).