The Amphibulimidae (Mollusca: Gastropoda: Orthalicoidea) from Colombia and adjacent areas
Author
Borrero, Francisco J.
Author
Breure, Abraham S. H.
text
Zootaxa
2011
3054
1
59
journal article
46210
10.5281/zenodo.278895
3498da36-33bd-4135-a475-9c978c7f7abc
1175-5326
278895
Dryptus pardalis
(Férussac, 1821)
(
Figs 2
A–B, 4A)
Helix pardalis
Férussac 1821: 48
.
Type
locality: not given.
Plekocheilus pardalis
(Férussac)
;
Tello 1975
: 201
.
Dryptus pardalis
(Férussac)
;
Breure 1984
: 490
, 495;
Richardson 1995
: 201
(references).
Bulimus leptochilus
Pfeiffer 1848b
: 48
.
Type
locality: La Baja, Prov. Pamplona.
Bulimus lindeni
Reeve 1848
: pl. 31 fig. 189.
Type
locality: La Baja, Prov. Pamplona.
Bulimus plumbeus
Pfeiffer 1855b
: 114
.
Type
locality:
Venezuela
.
Lectotype
BMNH 1975130 (
Breure 1978: 31, pl. 11 fig. 1
).
Material.
Cesar, foothills of Sierra de Perijá,
10–30 km
E of Becerril, El Roncón, B. Malkin leg./
1968–1969
:
FMNH
163692–163694; Socorpa Mission trail [approx.
09° 47’ N
73° 02’ W
,
1350 m
; see Harvey
et al.
2008], Becerril, Sierra de Perijá, Malkin leg./1968:
FMNH
159207; Guajira, [Arroyo] Potrero de Venancio,
1100 m
[not seen] (
Breure 1984
); hills S of Carraipía, O.L. Haught leg./1943:
USNM
599500; near Carraipía, Haught leg.:
USNM
543494; vicinity of Carraipía, Haught leg./1947:
MCZ
183880; Norte de Santander, Cúcuta, Hno. Nicéforo María leg./1936:
ANSP
168419; Hacienda Villa Felisa,
18 km
S of Cúcuta,
1100 ft
[~
333 m
, erroneous; actual elevation approx.
560 m
], M.A. Carriker leg./1947:
USNM
590659; Palogordo, Táchira Valley, S of Cúcuta,
4500– 5500 ft
[~
1360–1660 m
], Carriker leg./1947:
USNM
590635; Petrolia [sic, = Petrólea], Olsson leg./1935:
ANSP
166132; Petrólea, near sea level, Carriker leg.:
USNM
517813; Ocaña, G.
Wallis
leg./1875:
ZMUZ
511536; Buenos Aires, Highway Abrego–Sardinata,
6000–7000 ft
[~
1800–2120 m
], Carriker leg./1947:
USNM
486845; “Sta Librada” [= Santa Librada] (Sarare), Rochereau leg./1937:
DMNH
141725; Santander, La Baja (
Baker
1926
).?,
New Granada
:
CMC
C11024.
Venezuela
, Táchira, La Fría, H.B.
Baker
leg.:
UMMZ
244280;?,
120 km
E of Pamplona, Hno. Nicéforo María leg.:
ANSP
170736
.
Altitudinal range.
The localities reported here range between “near sea level” and
2435 m
.
Remarks.
Bulimus leptochilus
Pfeiffer, 1848
and
Bulimus lindeni
Reeve, 1848
were placed in the synonymy of this species by
Pilsbry (1895: 40)
. The
type
locality for both is “La Baja, Province of Pamplona.”
Richardson (1995: 202)
was the first to group
Bulimus plumbeus
Pfeiffer, 1855
, hitherto considered a species of
Thaumastus
Albers, 1860
, together with
Dryptus pardalis
. See also the remarks under
D. moritzianus
, some records of which might be misidentified given that these species are closely related. There seem to be two distinct shapes of what is referred to as
Dryptus pardalis
: (1) some are cigar-shaped, tapering about evenly at either side, with the central part having near-parallel outlines; and (2) some with the spire substantially less wide, and the body whorl wider, so the whole shell is more “bulimuliform” in appearance. In addition, there is substantial variation in sculpture, with some specimens being nearly smooth (even fresh ones), and others are very sculptured.
Two samples are tentatively referred to this species. One is from near Cúcuta (UVZ 8101), corresponding to the second shape variation above. The other is a lot from Guajira, near Carraipía, (USNM 543492). While these specimens look very different from other
Dryptus
, they might be misshapen
D. pardalis
. Specifically, they are too short, and have a very globose last whorl, which includes a much shorter aperture. One of the specimens has a strong parietal callus. Two reasons make it probable that they are
D. pardalis
: (1) this species is known from the same locality (it is the only
Dryptus
known from Guajira); (2) two other lots of
D. pardalis
are known from Carraipía: USNM 543494 (
2 specimens
) and USNM 599500 (
1 specimen
). Furthermore, one could argue that the deformed specimens are shorter and globose due to climatic differences. Interestingly, the spire is not globose, it seems to be just the body whorl having this characteristic. However, the following arguments are against this classification: (1) two specimens are available and both are somewhat similarly misshapen; (2) these individuals are umbilicate (more so than any specimen of
D. pardalis
); (3) neither one has much of a columellar fold. As is the case for several other species of
Dryptus
,
D. pardalis
may be widely distributed in
Venezuela
;
Tello (1975)
listed four Venezuelan Estados (states) in the distribution of this species, including Táchira and Zulia (bordering with
Colombia
), with Cumaná (Sucre) being the easternmost record. However, these records are in need of critical review.