Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae) Author Santos, Daubian https://orcid.org/0000-0003-1220-1267 Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil daubians@gmail.com Author Santos, Rodrigo dos Reis Pos-Graduacao em Entomologia, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Parana, Brazil Author Ribeiro, Guilherme Cunha Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil text Arthropod Systematics & amp; Phylogeny 2022 2022-09-12 80 439 494 http://dx.doi.org/10.3897/asp.80.e83035 journal article http://dx.doi.org/10.3897/asp.80.e83035 1864-8312-80-439 62FFB94CEBF441639F22881435EFC37C 3F4E21B524465471831D09738C2304A6 3.3.20. Amphineurus (Rhamphoneurus) oosterbroeki sp. nov. Material examined. Holotype : ♂ , Chile , Chiloe Is. , Chaiten [ 42°58′S 72°32′W ], 5/ 8-II-1954 , Pena (USNM) *. Paratypes : 1 ♂ , Chile , Chiloe Is. , Chaiten [ 42°58′S 72°32′W ], 5/ 8-II-1954 , Pena (USNM) *; 1 ♂ , Chile , Llanquihue , Carelmapu [ 41°44′S 73°40′W ], Coast of Canal de Chacao , 18-III-1957 , Pena (USNM) *; 1 ♂ , Chile , Osorno , Bahia Mansa [ 40°35′S 73°44′W ], South of Pucotrihue , 12-IV-1968 , Pena (USNM) *. Etymology. The species is named after Pjotr Oosterbroek, an important Tipulomorpha expert. Diagnosis. This species is distinguished by only a dark spot on R2, basal deflection of R5 as long as r-m, and M1+2 longer than basal deflection of M1. Furthermore, this species is characterized by male terminalia with sheath of aedeagus with a triangular spur and the tip of sheath inclined. Description. Wing length 5.47 mm, width 1.68 mm. - Coloration : General coloration brown. Rostrum and palpus brown. First segments of antenna yellow, remainding segments yellowish brown. Head brown. Abdomen yellowish brown. - Head (dorsal view Fig. 25B , ventral view Fig. 25A ): Rostrum bifurcated with long appendices; first palpal segment shorter than terminal segment; scape narrower than pedicel; terminal flagellomere dilated. - Thorax : Wing (Fig. 25D ) almost clear of markings, except for marking on R2; veins near fork of bM (Fig. 25C ) faded; M1+2 2.5 x longer than basal deflection of M1; R2+3+4 longer than R2+3. - Male terminalia (Fig. 25E ): Male tergite IX slightly darkened along posterior margin, with deep median U-shaped notch. Ventral branch of gonocoxite slender, very long, setose on distal tip. Lobe of gonostylus with rounded lobule; lobule longer than wide, shorter than half of lobe of gonostylus, narrower than stem. Clasper of gonostylus with differently shaped branches: lateral branch straight; medial branch curved. Mesal lobes of gonocoxite asymmetrical; left mesal lobe curved; right mesal lobe short with sharp turn. Sheath of aedeagus straightened, with wide rectangular projection; darkened only near tip. Figure 25. Amphineurus (Rhamphoneurus) oosterbroeki sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E Male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh , sheath of aedeagus; cgonst , clasper of gonostylus; goncx , gonocoxite; lgonst , lobe of gonostylus; ms lobe , mesal lobe of gonocoxite; t9 , male tergite IX. Remarks. Specimens of this new species were previously identified by C.P. Alexander as A. (R.) insanus Alexander. This species resembles A. (R.) podenasi sp. nov. but differs mainly in the length of r-m, concentration of setae on gonocoxite, shape of male tergite IX and right mesal lobe.