Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae) Author Santos, Daubian https://orcid.org/0000-0003-1220-1267 Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil daubians@gmail.com Author Santos, Rodrigo dos Reis Pos-Graduacao em Entomologia, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Parana, Brazil Author Ribeiro, Guilherme Cunha Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil text Arthropod Systematics & amp; Phylogeny 2022 2022-09-12 80 439 494 http://dx.doi.org/10.3897/asp.80.e83035 journal article http://dx.doi.org/10.3897/asp.80.e83035 1864-8312-80-439 62FFB94CEBF441639F22881435EFC37C 3F4E21B524465471831D09738C2304A6 3.3.21. Amphineurus (Rhamphoneurus) podenasi sp. nov. Material examined. Holotype . ♂ , Chile , Arauco , Nahuelbuta , Pichinahuel [ 37°48′S 73°02′W ], 1400-1600 m , 12/ 20-II-1953 , Pena (USNM) *. Paratypes : 1 ♂ , Chile , Chiloe Is. , Rio Coluco [ 42°6′S 73°55′W ], 30-I-1952 , Pena (USNM) *; 1 ♂ , Chile , Llanquihue , Carelmapu [ 41°44′S 73°40′W ], Coast North of Canal de Chacao , 18-III-1957 , Pena (USNM) *. - Additional material : CHILE . 1 ♀ and 1 [sex unknown], Chiloe Is. , Dalcahue [ 42°22′S 73°39′W ], II-1954 , Pena (USNM); 1 [sex unknown], Chiloe Is. , Aulen [ 42°2′S 74°1′W ], 8-II-1952 , L.P. Guzman (USNM); 1 [sex unknown], Chiloe Is. , Chaiten [ 42°58′S 72°32′W ], 5/ 8-II-1954 , Pena (USNM) . Etymology. The species is named after Sigitas Podenas, a skilled Tipulomorpha expert. Diagnosis. This species is distinguished by only a dark spot on R2 and basal deflection of R5 longer than r-m. Furthermore, this species is characterized by male terminalia with sheath of aedeagus with a spur, tip of right mesal lobe curved downwards and sheath of aedeagus with rounded spur. Description. Wing length 5.73 mm, width 1.83 mm. - Coloration : General coloration brownish-yellow. Rostrum and palpus brown. First segment of antenna yellow, remainding segmens dark yellow. Head dark brown. Eyes black. Thorax brownish yellow, pleura pale. Contrasting golden setae on prescutum. Scutum without stripes. Halter brownish-yellow. Coxae and legs brownish yellow. Abdomen dark brown. - Head (dorsal view Fig. 26B , ventral view Fig. 26A ): Rostrum bifurcated with long appendices; first palpal segment shorter than terminal segment; scape thinner than pedicel; flagellomeres gradually reducing in length, resulting in shorter terminal flagellomeres. - Thorax (lateral view Fig. 26E , dorsal view Fig. 26F ): Thorax with several setae, including long setae on prescutum. Anatergite shorter than katatergite. Halter with dilated knob. Wing (Fig. 26D ) almost clear of markings, except for strong marking covering R2; veins near fork of bM (Fig. 26C ) faded; M1+2 longer than basal deflection of M1; R2+3+4 shorter than both R2+3 and basal deflection of R5. - Female terminalia (Fig. 26H ): Female tergite IX longer and wider than tergite X; cercus shorter than hypoginial valves, curved dorsad. Both cercus and hypogynial valve very slender. - Male terminalia (Fig. 26G ): Male tergite IX slightly darkened along posterior margin, with wide V-shaped median notch. Ventral branch of gonocoxite inclined medially in distal tip. Dorsal branch of gonocoxite nearly half-length of ventral branch. Lobe of gonostylus with rounded lobule; lobule longer than wide, longer than half-length of lobe of gonostylus, narrower than stem. Clasper of gonostylus with differently shaped branches: lateral branch straight; medial branch curved, club shaped. Mesal lobes of gonocoxite asymmetrical; left mesal lobe short with sideways projection; right mesal lobe curved. Sheath of aedeagus straightened, with rounded projection; darkened only on tip. Figure 26. Amphineurus (Rhamphoneurus) podenasi sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh , sheath of aedeagus; anatg , anatergite; anepm , anepimeron; anepst , anepisternum; aprn , anteropronotum; cer , cercus; cgonst , clasper of gonostylus; cx , coxa; goncx , gonocoxite; hlt , halter; hyp val , hypogynial valve; k , knob of halter; kepm , katepimeron; kepst , katepisternum; ktg , katatergite; l ms , left mesal lobe of gonocoxite; lgonst ; lobe of gonostylus; mr , meron; ms lobe , mesal lobe of gonocoxite; mtanepst , metanepisternum; mtepm , metepimeron; mtg , mediotergite; mtkepst , metakatepisternum; mtn , metanotum; p , posterior basalare; patg , paratergite; pprn , postpronotum; presct , prescutum; r ms , right mesal lobe of gonocoxite; sct , scutum; sctl , scutellum; st , sternite; tg , tergite; t9 , male tergite IX. Remarks. Specimens of this new species were previously identified by C.P. Alexander as A. (R.) insanus Alexander. This species resembles A. (R.) oosterbroeki sp. nov. but differs mainly in the length of r-m, concentration of setae on the gonocoxite, shape of male tergite IX and right mesal lobe.