A Catalogue and Historical Study of the Odonata of Ecuador
Author
Mauffray, William F.
Author
Tennessen, Kenneth J.
text
Zootaxa
2019
2019-07-03
4628
1
1
265
journal article
26210
10.11646/zootaxa.4628.1.1
3e77e5cf-7e8a-490d-a5f7-c571b90aea57
1175-5326
3335707
E4C2F52F-9A1A-4C55-B03B-C87B2B4C4676
Argia
Rambur, 1842
Argia
is the largest genus of damselflies in the New World, with over 138 valid species names and at least seven species yet to described (
Garrison & von Ellenrieder 2018
). Adults range from small to large and are variable in color pattern, the thoracic dorsum being black with blue or purple stripes (rarely orange) to entirely metallic red or copper, the thoracic sides usually blue, and the abdomen from mostly blue with black markings to mostly black with small blue markings (
Figs. 16a & 16b
), rarely largely orange. Species identification relies on Calvert (1902), and
Garrison & von Ellenrieder (2015
, 2017, 2018). Thirty-two species have been recorded in
Ecuador
, of which three are considered endemic. Habitat of most species in
Ecuador
is small streams to mid-sized rivers, with a few found in pooled areas of slow streams, seepage streams or sheet flow over boulders.
FIGURE 15.
Anisagrion inornatum
♂
,
ECUADOR
,
Loja Province
., seep, trib. of River Chamba, Vilcabamba,
30 iii 2008
; photo K. J. Tennessen.
Argia acridens
Garrison & von Ellenrieder, 2018
. Publ. Recs.
ES:
*
Garrison & von Ellenrieder (2018)
;
MN:
*
Garrison & von Ellenrieder (2018
—
Holotype
: Flavio-Alfaro Canton:
79km
west of Santo Domingo de los Colorados (
-0.3333
,
-79.6666
,
260m
),
7 v 1975
, leg. Paul J. Spangler
et al.
[USNM]);
SD: *
Garrison & von Ellenrieder (2018)
.
Note: Endemic to Ecuador’s Pacific slope.
Argia adamsi
Calvert, 1902
. Publ. Recs.
ECU: (
P11), (H14);
LR
:
Garrison & von Ellenrieder (2018)
;
MN:
Garrison & von Ellenrieder (2018)
;
MS:
Garrison & von Ellenrieder (2018)
;
NA:
*
Tennessen (1996b)
, Daigle (1996),
Garrison & von Ellenrieder (2018)
;
OR:
Garrison & von Ellenrieder (2018)
;
PA:
Garrison & von Ellenrieder (2018)
;
PC:
Garrison & von Ellenrieder (2018)
;
SD:
Garrison & von Ellenrieder (2018)
;
TU:
Garrison & von Ellenrieder (2018)
. Also CEN, PER, VEN: (H08), (P11); Plus COL: (H08), P&P (2011), (P16), and BOL, BRA:
Garrison & von Ellenrieder (2018)
.
Addl. Data.
ES
,
San Lorenzo:
[FSCA. JJD, KJT];
LR
,
Quevedo:
[FSCA, JTJ, KJT];
MS
,
Gualaquiza:
[USNM];
Huambaya:
[KJT, RWG];
NA
,
Tena:
[FAL, FSCA, UTIC, JJD, KJT, RWG, TWD, USNM];
OR
,
Francisco de
Orellana
:
[FSCA, JJD, KJT, RWG, SWD];
Loreto
[FSCA, JTJ, KJT, RWG, USNM];
PA
,
Pastaza
:
[FSCA, JJD, JTJ, KJT, RWG, USNM];
SD:
[RWG];
SU
,
Lago Agrio:
[SWD];
Shushufindi:
[UTIC].
Note: In east slope form the light coloration on the thorax is blue, and in some west slope forms it is lighter blue or pale yellowish, especially from northwest
Ecuador
. According to
Garrison & von Ellenrieder (2018)
, L. K. Gloyd considered
A. adamsi
from
Ecuador
west of the Andes to represent a new species to which she had applied the manuscript name
A. lorena
. However,
Garrison & von Ellenrieder (2018)
found no significant differences in male caudal appendages nor female mesostigmal plates.
FIGURE 16a.
Argia fulgida
♂, ECUADOR, Pichincha Province, small stream, Mashpi Road, 2 x 2018; photo K. J. Tennessen.
FIGURE 16b
.
Argia pulla
♂, ECUADOR, Bolivar Province., stream 5 km E of Caluma, 5 iii 2010; photo K. J. Tennessen.
Argia collata
Selys, 1865
. Publ. Recs.
ECU
: *Tennessen (1996b—as
A. thespis
), (L06—as
A. thespis
), (H08), (H14),
Garrison & von Ellenrieder (2018)
;
NA:
*
Garrison & von Ellenrieder (2015)
;
OR:
*
Garrison & von Ellenrieder (2015)
;
SU: *
Garrison and von Ellenrieder (2015)
. Also BRA: (L06—as
A. thespis
),
Garrison & von Ellenrieder (2015)
, (H08), (P11—as
A. thespis
), GUF: (H08); plus COL, PER, VEN:
Garrison & von Ellenrieder (2015)
; and BOL (P16).
Addl. Data.
EO
,
Santa Rosa
:
1 ♂
, stream
12 km
E of Arenillas, Hwy. 25 (
-3.5244
,
-79.9831
,
30m
),
29 iii 2008
, leg. & det. KJT—as
A. thespis
[FSCA];
GY
,
Balao
:
1 ♂
, stream, banana plantation,
2 km
E of Hwy. 25 & Balao Jct. (
-2.9100
,
-79.6919
,
70m
),
28 iii 2008
, leg. & det. KJT—as
A. thespis
, [FSCA];
NA, Tena
:
4 ♂♂
,
1 ♀
, Río Chinchipino,
27.7 km
E of Puerto
Napo
(
-1.0728
,
-77.5864
,
385m
),
13 xi 1997
, leg. & det. J. Daigle—as
A. thespis
[JJD];
OR
,
Francisco
de
Orellana
:
1 ♂
, stream
2.3 km
E of Yasuni Sci. Res. Sta.
, P.N.Y.,
9 vi 1995
, leg. & det. KJT—as
A. thespis
[KJT];
1 ♂
, same data but leg. & det. WFM—as
A. thespis
[FSCA]
;
1 ♂
, seepage
2.1 km
E of first road past research station,
Yasuni
Preserve,
15 xi 1997
, leg. & det. JJD—as
A. thespis
[JJD];
1 ♀
, stream
Waorani
Road
, 2.0 km past cutoff,
Yasuni
Preserve
,
16 xi 1997
, leg. & det. JJD—as
A. thespis
[JJD];
SU
,
Cascales
:
1 ♀
, stream
50 km
W of Lago Agrío
(
395m
),
8 vi 1995
, leg. & det. KJT—as
A. thespis
[KJT];
Shushufindi
:
1 ♂
, stream about
3 km
NW of Shushufindi
(
-0.1664
,
-76.6167
,
280m
),
14 v 2009
, leg. & det. KJT—as
A. thespis
[KJT].
Note: According to Garrison (pers. comm. 2011), “
A. collata
was thought to be a synonym of
A. thespis
, but it has been elevated back to distinct species. Specimens labeled and/or published as
A. thespis
from
Ecuador
should be this species.”
Argia cuspidata
Garrison & von Ellenrieder, 2018
. Publ. Recs.
ES:
*
Garrison & von Ellenrieder (2018
—
♀
para- type, from tandem pair),
San Lorenzo:
seeps
1.1 km
NW of Lita at bridge, 5:00 pm [
0.8850
,
-78.4755
,
470m
],
2 ii 1997
, leg JJD, [JJD]);
SD:
*
Garrison & von Ellenrieder (2018
—
Holotype
:
19 km
E of Santo Domingo de los Colorados [
-0.3137
,
-79.0289
,
740m
],
7 v 1975
, leg. A. Langley & J. Cohen [USNM]).
Note: Endemic to limited area of Pacific slope.
Argia difficilis
Selys, 1965
. Publ. Recs. ECU: (H08); (P11);
MS:
Garrison & von Ellenrieder (2007);
NA:
Garrison & von Ellenrieder (2007);
OR:
Garrison & von Ellenrieder (2007);
PA
: Garrison & von Ellenrieder (2007);
SU:
Garrison & von Ellenrieder (2007). Also BRA, CEN, COL, PER, VEN, T&T: (H08).
Addl. Data.
MS
,
Gualaquiza:
[RWG, UMMZ, USNM];
NA, Tena:
[DRP, FSCA, RWG, UMMZ];
OR, Francisco de
Orellana
:
[FSCA, KJT, NVE, SWD, TWD, USFQ, USNM];
Loreto
:
[KJT, RWG, UMMZ];
PA, Mera:
[FSCA];
Pastaza
: [RWG];
SU
,
Shushufindi
: [DRP, FSCA, RWG, SWD].
Argia dives
Förster, 1914
. Publ. Recs.
ECU:
(L06), (H08), (P11), (H14);
MS
: *
Tennessen (1998)
,
Garrison & von Ellenrieder (2018)
;
NA:
Garrison & von Ellenrieder (2018)
;
OR:
Garrison & von Ellenrieder (2018)
;
PA
:
Tennessen (2006)
,
Garrison & von Ellenrieder (2018)
;
SU:
(
Garrison & von Ellenrieder, 2018
);
TU:
Garrison & von Ellenrieder (2018)
. Also PER: (L06), (P11); plus BRA: (L06); and COL: P&P (2011), (P16).
Addl. Data.
MS, Huamboya:
[JJD];
Morona:
[RWG];
Gualaquiza:
[FSCA];
NA
,
Archidona:
[RWG];
Carlos Julio Arsemena Tola
: [RWG];
Tena:
[FSCA, KJT, RWG, USNM];
OR, La Joya de los Sachas:
[FSCA];
Loreto
:
[DRP, FSCA, JJD, JTJ, KJT, RWG, TWD, USNM];
PA
,
Mera:
[FSCA, JJD, KJT, RWG, USNM];
Pastaza
:
[FAL, FSCA, JJD, KJT, RWG, SWD, SWD];
SU
,
Shushufindi:
[FSCA, RWG, SWD];
ZC
,
El Pangui:
[FSCA, JJD, KJT].
Argia euphorbia
Fraser, 1946
. Publ. Recs.
ECU:
(P16),
Garrison & von Ellenrieder (2018)
; *
OR
:
von Ellenrieder & Garrison (2011)
,
Garrison & von Ellenrieder (2015)
.
Addl. Data.
OR
,
Francisco
de
Orellana
:
1 ♂
, T.
B.S.
(
-0.6333
,
-76.1500
,
220m
), date?, leg.
K.
Jackson
,
D. Inward
, det.
RWG-2003
[BMNH]
Also
PER: (
H08
), (P11); plus ARG, BRA: (
H08
); plus COL, VEN: (
P16
).
Note: The
Argentina
record is an error (
Garrison & von Ellenrieder 2015
).
Argia fraudatricula
Förster, 1914
. Publ. Recs.
ECU:
(*H08), (P11), (H14),
Garrison & von Ellenrieder (2018)
. Also PER: (H08), (P11).
Addl. Data.
NA
,
Tena
:
1 ♂
, Jatun Yacu, near mouth of Ilapulin,
8 iii 1937
, leg. WCM, det. JJD [FSCA]; same but
4 ♂♂
, det.
L. Gloyd
, from RWG [RWG, UMMZ];
3 ♂♂
,
Jatun Yacu
,
Río
Napo
watershed (
-1.0167
,
-77.8332
,
700m
),
4 iii 1937
, leg.WCM, det.
L. Gloyd
, from UMMZ [RWG, UMMZ]; same but
3♂♂
,
2 ♀♀
,
21 iii 1937
, [RWG, UMMZ];
PA
:
4 ♂♂
,
Partidero River
trail [
El Partidero
, Indian village on east bank of Río Anzu, jct. of trails from Mera and Puyo northward to
Napo
],
Río
Napo
watershed (
-1.0167
,
-78.000
,
1000m
),
16 ix 1935
, leg. WCM, det.
L. Gloyd
, from RWG [RWG, UMMZ];
ZC
,
Nangaritza
:
7 ♂♂
,
2 ♀♀
,
Río Nangaritza
(-4.55, -78.8167,
1400m
),
2 xii 1941
, leg.
D. Laddey
, det.
L. Gloyd
[RWG, UMNZ]; same but
1♂
,
1 ♀
,
Mayaicu
,
Río Nangaritza
(
- 3.9894
,
-78.6328
,
1136 m
) [RWG]; same but
2 v 1941
[RWG].
Note: There are no recent records of this species despite numerous field surveys within that part of
Ecuador
where it was recorded more than 75 years prior to this report.
Argia fulgida
Navás, 1934
. Publ. Recs.
BO:
*
Garrison & von Ellenrieder (2017
,
2018
);
EO:
*
Garrison & von Ellenrieder (2017
,
2018
);
ES:
*
Garrison & von Ellenrieder (2017
,
2018
);
LR:
*
Garrison & von Ellenrieder (2017
,
2018
);
MN:
*
Garrison & von Ellenrieder (2017
,
2018
);
PC:
Garrison & von Ellenrieder (2018)
;
SD:
*
Garrison & von Ellenrieder (2017
,
2018
);. Also CEN, COL:
Garrison & von Ellenrieder (2017)
.
Addl. Data.
BO, Caluma:
4 ♂♂
, rocky stream,
5 km
E of Caluma, Guaranda Road (
-1.5942
,
-79.2308
,
480m
),
5 iii 2010
, leg. JJD, det. RWG [JJD, RWG];
4 ♂♂
, same data but KJT [FSCA, KJT];
CN,
Cañar
:
2 ♂♂
,
Cochancay
20.2 km
E of Prov. Border on Hwy.
70 rocky stream (
-2.4686
,
-79.2853
,
195m
),
3 xi 1997
, leg, det. JJD [FSCA];
EO,
Portovelo
:
1 ♂
,
Portovelo
(
-3.7167
,
-79.6167
,
650m
),
1 vii 1941
, leg.
D. Laddey
, det. RWG [RWG];
ES
,
San Lorenzo
:
1 ♂
, rocky stream
36.5 km
NW of Lita
, near Urbina,
3 ii 1997
, leg. & det. JJD; 1 tan. pair,
1 ♂
, [San Antonio], nearby,
36.5 km
WNW of Lita
on [old]
San Lorenzo Hwy
, pools/seeps/stream (0.95, -78.68,
200m
),
3 ii 1997
, leg. & det. RWG [FSCA];
LR
,
Babahoyo
2 ♂♂
,
2 ♀♀
,
Playas de Juan Montalvo
, near
Balzapamba
(
-1.6833
,
-79.2833
,
15m
),
28 ii 1938
, leg. WCM, det. RWG, from UMMZ [RWG];
1 ♂
,
1 ♀
, same data but
8 iv 1938
[RWG];
2 ♂
, same data but
15 iv 1938
[RWG];
2 ♂♂
, same data but
17 iv 1938
[RWG];
1 ♂
, same data but
31 vii 1941
[RWG];
PC,
Quito
:
6 ♂♂
, upper Mashpi Road,
Rio Huaycuyacu
, above waterfall (
0.1952
,
-78.8657
,
850m
),
2 ix 2018
, leg. & det. WFM [FSCA, USFQ];
2 ♂♂
, 1 tan. pair, same data but leg & det. KJT [KJT];
SD:
3 ♂♂
,
1 ♀
,
Río Pove
,
Santo Domingo
de los
Colorados
(-0.25, -79.15,
600m
),
27 ix 1940
, leg.
D. Laddey
, det. RWG, from UMMZ [RWG];
1 ♂
,
1 ♀
, same data but
28 ix 1940
[RWG];
1 ♂
,
Río Mulaute
, near
Santo Domingo
de los
Colorados
; virgin forest,
3 iii 1973
, leg.
M. Deyrup
, det. RWG, from UMMZ [RWG];
1 ♂
, small stream,
Hwy.
30, near km marker 71, near
La Unión del Toachi
(
-0.3139
,
-78.9436
,
900m
),
25 ii 2010
, leg. & det. JTJ [JTJ].
Note: All records are from Pacific slope.
Argia fumigata
Hagen
in
Selys, 1865
. Publ. Recs.
ECU:
(P16),
Garrison & von Ellenrieder (2018)
; *
OR:
Garrison & von Ellenrieder (2015)
. Also: BRA, GUF, GUY, SUR, VEN: (P16).
Addl. Data.
OR:
Francisco
de
Orellana
:
1 ♀
,
Reserva Étnica
Waorani
,
1 km
S Okone Gare Camp
, canopy fogging project,
Lot
#1546, TR3 #6, terra firme forest (
-0.6569
,
-76.4528
,
216 m
),
21 vi 1996
, leg.
T.L. Erwin
et al
., det. RWG [USNM].
Note: This record is the westernmost for this predominately
Guiana
shield species.
Argia gerhardi
Calvert, 1909
. Publ. Recs. ECU: (H08), (P11), (H14). Also BOL, COL, PER: (H08), (P11).
Addl. Data. Provided by R. Garrison (pers. comm.,
25 vi 2018
):
LJ,
Loja
:
1 ♂
,
1 ♀
, Estación Cientifica San Francisco, Río San Francisco, small creek, oviposition in soil (
-3.9744
,
-79.0778
,
1870m
),
28 x 2014
, leg. C. Willigalla, det. RWG [
C. Willigalla
];
NA,
Archidona
:
4 ♂♂
,
1 ♀
, seepages near river,
43.5 km
W
Loreto
(
-0.7212
,
-77.6410
,
880m
),
14 vi 1995
, leg & det. SWD [RWG];
1 ♂
, seep
44 km
east of
Narupa
jct.,
Loreto
Road
,
14 xi 1997
, leg. KJT, det. RWG [RWG];
PA,
Mera
:
1 ♀
,
20 km
on
Puyo
to
Baños Hwy.
(
-1.4543
,
-78.1221
,
1130m
),
19 x 1997
, leg.
M. Kippenrand
, det. RWG [RWG].
SU,
Gonzalo Pizarro
:
1 ♂
,
1 ♀
,
San Rafael Falls
,
Río Quijos
,
52 km
northeast of
Baeza
(
-0.1020
,
-77.5890
,
1290m
),
22 i 2009
; leg DLW, WAH,
Michael Thomas
,
Fred Morrison
&
Ronald
Vargas
, det. RWG [WAH];
TU
,
Baños
:
1 ♀
,
Abitagua
,
Río
Pastaza
watershed,
Hda. Guillermina
(
-1.4431
,
-78.1665
,
1200m
),
7 xii 1939
, leg. WCM, det.
L. Gloyd
[RWG]; same but,
1 ♂
,
9 xii 1939
[RWG];
ZC,
Zamora
:
1 ♂
, seepage near
Río Sabanillo
,
W of Zamora
(
-4.0453
,
-79.0147
,
1225m
),
1 iv 2008
, leg. & det. JTJ [JTJ];
1 ♂
,
1 ♀
, tributary of
Bombuscara River
on road towards
Zamora
(
-4.0428
,
-79.0142
,
1280m
),
18 viii 2006
, leg. JCA, det. RWG [UTIC].
Notes: According to Rosser Garrison (pers. comm.
25-vi-2018
), “The identification within the
Argia gerhardi
/
kokama
/
nigrior
complex is difficult. I have seen (and illustrated)
types
of all three and have thus far done a lot of work on this group....Putting names on various populations is not always easy as I have had to compare various characters from Ecuadorean and Peruvian populations with the
types
(
Bolivia
,
Peru
) and the characters I have thus far seen to be of diagnostic value do not seem to be concordant. In particular, males of
Argia kokama
approach those of
A. nigrior
closely and the females of
A. gerhardi
and
A. nigrior
are difficult to separate. All of this is going to require more work.....the records I have for
Argia gerhardi
from
Ecuador
are listed with the proviso that not all of the records may pertain to that species.”
All records are from relatively high elevation, from
880–1870 m
.
Argia hamulata
Fraser, 1946
. Publ. Recs. ECU: *
Garrison & von Ellenrieder (2018)
, Also BOL (RWG, pers. comm.), PER: (H08). So far known from only one record:
SD:
1 tan. pair, stream, Tinalandia, E of Santo Domingo, Hwy. 30 (
-0.3000
,
-79.0667
,
800m
),
31 i 1997
, leg. KJT, det. RWG, data provided by RWG [KJT].
Argia huanacina
Förster, 1914
. Publ. Recs.
ZC:
stream, north side of Guaguayme Bajo,
4 iv 2008
, leg. & det. KJT [FSCA] *
Tennessen & Johnson (2008)
,
Garrison & von Ellenrieder (2018)
. Also BOL, PER (H08) (P11),
Garrison & von Ellenrieder (2018)
.
Notes: From Rosser Garrison (pers. comm., 2018): “This species was treated and illustrated in
Garrison & von Ellenrieder (2015)
. von Ellenrieder &
Garrison (2007)
followed by
Heckman (2008)
illustrated the appendages of a male in the RBINS with the manuscript name “
lateralis
” by Selys as
A. huanacina
. This is incorrect. Subsequent examination of these specimens showed the manuscript name
lateralis
Selys
to represent
A. infrequentula
as can be seen by comparing
Figs. 6
l-n in von Ellenrieder &
Garrison (2007)
and Fig. 3.1.
427 in
Heckman (2008)
to
Figs. 10
l-2 and 10 p-
2 in
Garrison & von Ellenrieder (2015)
.”
This southern
Ecuador
record seems to be the northernmost record for this species.
Argia inculta
Hagen
in
Selys, 1865
. Publ. Recs.
ECU:
(*H08), (P11), (H14),
Garrison & von Ellenrieder (2018)
. Also PER: (H08), (P11).
Addl. Data.
ZC
:
2 ♂♂
,
1 ♀
, Pacocha (
-1.0577
,
-80.8472
,
120m
),
27 iv 1946
, leg. L. Gomez Alonzo, det. L. Gloyd [RWG, UMMZ];
EO
,
Arenillas
: 1 tan. pair,
2 ♂
, clay-bottom stream,
14 km
S of Arenillas, road to Alamar (
-3.6656
,
-80.0869
,
95m
),
10 iv 2008
, leg. & det. KJT [FSCA];
3 ♂♂
, same data but leg. & det. JTJ [JTJ]; 1 tan pair, farm stream,
11 km
S of Arenillas, road to Alamar (
-3.6436
,
-80.0989
),
10 iv 2008
, leg. & det. KJT [FSCA];
Piñas
:
1 ♀
, Piñas (-3.6667, -79.65,
1000m
),
23 vii 1941
, leg. D. Laddey, det. L. Gloyd, from CHK/ UMMZ [RWG];
Portoviejo:
9 ♂♂
,
2 ♀♀
, Cerro de Hojas (
-1.0432
,
-80.4311
,
200m
),
19 iv 1946
, leg. L. Gomez Alonzo, det. MJW [CSCA, RWG, UMMZ], same but,
21 iv 1946
,
10 ♂♂
,
3 ♀♀
, [CSCA, RWG, UMMZ]; same but,
16 ♂♂
,
2 ♀♀
, [CSXA, RWG, UMMZ];
1 ♀
, concrete ditch, Hwy. E40, E of Portoviejo (
-1.0672
,
-80.1350
,
420m
),
4 iii 2010
, leg. & det. KJT [FSCA];
Santa Rosa
:
1 ♂
, stream,
12 km
E of Arenillas, Hwy. 25 (
-3.5244
,
-79.9831
,
30m
),
29 iii 2008
, leg. & det. KJT [FSCA];
2 ♂♂
, same data but leg. & det. JTJ [JTJ];
2 ♂♂
, same data but
9 iv 2008
, leg. & det. KJT [FSCA];
LJ, Catamayo:
1 ♂
,
1 ♀
, small stream and tilapia pond on old road from Malacatos to Catamayo, (
-4.0312
,
-79.3558
,
1290 m
),
18 i 2014
, leg. F. Morrison, R.
Vargas
, & E. Wyman, det. RWG [RWG];
MN
,
Chone
:
2 ♂♂
,
7.2 km
NNE of Chone, Boca de Mosquito, between Río Grande and Río Mosquito (
-0.6833
,
-80.1000
,
110m
),
24 x 1978
, leg. J. Anderson, det. RWG, from USNM [JJD, RWG];
1 ♂
, small stream/trib.
3 km
N of Canuto
(
-0.7678
,
-80.1231
,
60m
),
3 iii 10
, leg. & det. JTJ [JTJ];
9 ♂♂
, same data but leg. & det. JJD [JJD];
Flavio Alfaro
:
1 ♂
, borrow pit/small stream about
4 km
N of Flavio Alfaro
(
-0.3750
,
-79.8875
,
295m
),
3 iii 2010
, leg. & det. JTJ [JTJ];
Jipijapa:
1 ♂
, [Playa]
San Jose
,
4 miles
inland (-1.38, -80.75,
60m
),
12 iii 1970
, leg.? det. KJT [FSCA];
2 ♂♂
, Puerto Lopez,
20 km
to Salinas,
5 iv 1998
, leg.
E. Tapia
, det. JJD [JJD];
Junin
:
2 ♂♂
,
1 ♀
, stream just S of
Junin
(
-0.9317
,
-80.2256
),
3 iii 2010
, leg. & det. KJT [FSCA];
Manta:
1 ♂
, Santa Rosa (
-1.0576
,
-80.8472
,
20m
),
11 v 1946
, leg.
L. Gomez Alonzo
, det.
L. Gloyd
[RWG]; same but,
6 ♂♂
,
2 ♀♀
,
12 v 1946
[CSCA, RWG, UMMZ]; same but,
6 ♂♂
,
1 ♀
,
14 v 1946
[CSCA, RWG, UMMZ];
Pedernales
:
2 ♂
, small stream about
2 km
E of Pedernales
(
0.0836
,
-80.0139
,
90m
),
28 ii 2010
, leg. & det. KJT [FSCA];
1 ♂
, same data but leg. & det. JTJ [JTJ].
Note: Collection data indicate that
A. inculta
is a low elevation west slope species.
Argia indicatrix
Calvert, 1902
. Publ. Recs.
ECU:
(L06), (H08), (P11), (H14),
Garrison & von Ellenrieder (2018)
;
CN:
Garrison & von Ellenrieder (2015)
;
EO:
Garrison & von Ellenrieder (2015)
; *
ES:
Daigle
et al.
(1997)
,
Garrison & von Ellenrieder (2015)
;
LR:
Garrison & von Ellenrieder (2015)
;
NA:
Garrison & von Ellenrieder (2015)
;
OR:
von Ellenrieder & Garrison (2011)
,
Garrison & von Ellenrieder (2015)
;
PC:
Garrison & von Ellenrieder (2015)
; *
SD:
Daigle
et al.
(1997)
Garrison & von Ellenrieder (2015)
;
SU:
Garrison & von Ellenrieder (2015)
. Also CEN, COL, PER, VEN: (L06), (H08), (P11) plus BRA: (L06), (H08) and BOL: (P16).
Addl. Data:
BO, San Miguel:
[RWG];
CN, La Troncal:
[RWG, SWD];
EO, Santa Rosa:
[FSCA, JTJ, KJT];
ES, San Lorenzo:
[FAL, FSCA, JA, JJD, KJT, RWG, SWD];
LR, Babahoyo:
[RWG];
Buena Fé:
[DRP, SWD];
MN, Flavio Alfaro:
[FSCA, KJT];
OR, Francisco de
Orellana
:
[JJD, KJT, RWG, SWD];
Loreto
:
[JTJ, KJT];
PC, San Miguel Bancos:
[RWG]; SD: [DRP, FAL, FSCA, JJD, KJT, RWG, SWD];
SU, Lago Agrío:
[RWG].
Argia infrequentula
Fraser, 1946
. Publ. Recs.
ECU:
(H08), (P11), (H14),
Garrison & von Ellenrieder (2018)
;
NA
: *Daigle (1996). Also COL: (H08, P11, P&P); and PER: (P16).
Addl. Data.
MS, Morona
: [UMMZ];
NA, Tena:
[BMNH, CSCA, FSCA, JJD, KJT, RWG, UMMZ];
OR, Francisco de
Orellana
:
[FSCA, KJT, TWD, USNM];
Loreto
:
[JJD, KJT, USNM];
PA, Mera:
[JJD, KJT, RWG, UMMZ];
Pastaza
:
[BMNH, CSCA, RWG, UMMZ];
Santa Clara:
[JJD, RWG];
SU, Cascales:
[FSCA, USNM];
Lago Agrio:
[RWG];
Sushufindi:
[FAL, FSCA, JJD, JTJ, KJT, RWG];
TU, Baños
: [UMMZ];
ZC, Zamora:
[RWG].
Note: Some of these older records need to be rechecked as they may be
Argia selysi
Garrison & von Ellenrieder (2018)
.
Argia jocosa
Hagen
in
Selys, 1865
. Publ Recs. S
U:
1 ♂
, Bellavista River,
32km
SE of Lago Agrio,
11 i 2014
, leg. D. Wagner, W. Haber, F. Morrison, det. RWG [RWG]
Garrison & von Ellenrieder (2018)
. Also COL, VEN: (
Garrison & von Ellenrieder 2018
).
Argia kokama
Calvert, 1909
. Publ. Recs.
ECU:
Garrison & von Ellenrieder (2018)
;
MS
: *
Tennessen (2006)
;
OR:
von Ellenrieder & Garrison (2011
, 2017);
SU:
Garrison & von Ellenrieder (2017)
; Also PER: (H08), (P11); plus COL: P&P (2011), (P16).
Addl. Data.
MS, Huamboya:
6 ♂♂
, 1 tan. pair, Macas,
58 km
N on Hwy. 45 (
70 km
N of Sucua), roadside ditch and tribs. near Río Pasanac (
-1.9447
,
-77.8536
,
762m
),
16 ix 2005
, leg. & det. WFM [FSCA, QZAC, USNM];
3 ♂♂
,
1 ♀
, same but:
10 xi
197, leg. & det. JJD—as “
A. makoka
”
[JJD];
1 ♂
, same data but, leg. TWD, det. JJD—as “
A. makoka
” [JJD];
Morona
;
8 ♂♂
,
Macas
15.7 km
W on
Macas-Guamote Road
small stream (
-2.2689
,
-78.1936
,
1463m
),
17 ix 2005
, leg. & det. WFM [FSCA, QZAC, USNM];
8 ♂♂
,
Macas
22.9 km
W on
Guamote Road
,
3.3 km
W on
Río Albanico
rocky stream and flowing roadside ditches (
-2.2342
,
-78.218
, 1690m),
18 ix 2005
,
8 ♂♂
, leg. & det. WFM [FSCA, QZAC, USNM];
OR,
Francisco
de
Orellana
,
1 ♀
, T.
B.S.
, USFQ, open grassy area behind din- ing room, surrounded by bushes and trees and some seepage areas (
-0.6383
,
-76.1492
,
220m
),
14 i 2009
, leg. & det. RWG & NVE [RWG]; 2 tan. pairs, same data but
15 i 2009
; 1 tan. pair,
1 ♂
, same data but,
16 i 2009
,
1 ♂
, same data but
16 i 2009
,
1 ♂
,
1 ♀
, same data but
18 i 2009
;
1 ♂
, T.
B.S.
, USFQ, black water stream crossing
Guacamayo
trail at marker 2050,
16 i 2009
,
1 ♂
, same data but T.
B.S.
, USFQ,
Mauritia
palm swamp off
Guacamayo
trail at marker 2100,
20 i 2009
,
2 ♂♂
,
1 ♀
, same data but T.
B.S.
, USFQ,
Guacamayo
trail between markers 450–2100,
22 i 2009
, leg. & det. RWG & NVE [USFQ];
1 ♂
,
1 ♀
,
General Plaza Gutierrez
: [about
7 km
N] &
22.5 km
N of Indanza on Hwy.
45, roadside ditch,
10 i 2009
, leg. & det. RWG & NVE [RWG];
PA,
Santa Clara
:
El Partidero
,
Río
Napo
watershed (
-1.2833
,
-77.9000
,
800m
),
4 ix 1935
, leg. WCM, det. RWG, from C.H.
Kennedy
[UMMZ];
Mera
;
5 ♂♂
,
Mera
between
Mera
and
Shell
,
Los Copales
trails and forest
N of Río Metolo
(
-1.4886
,
-78.0708
,
1021m
),
20 ix 2005
, leg. det. WFM [FSCA, QZAC, USNM].
Argia limitata
Navás, 1924
. Publ. Recs. -NA: *
Garrison & von Ellenrieder (2018)
;
OR:
*Garrison & von Ellenrie- der (2018); PA: *
Garrison & von Ellenrieder (2018)
;
SU:
*
Garrison & von Ellenrieder (2018)
;
ZC
: *
Garrison & von Ellenrieder (2018)
. Also PER: (H08), (P11), (H14).
Addl. Data:
NA, El
Chaco
:
[FSCA];
Tena:
[JJD, KJT, RWG, SWD, SWD];
OR,
Loreto
:
[JTJ, KJT];
PA, Mera:
[JJD];
Pastaza
: [FAL, JJD, KJT, RWG, SWD];
SU, Cascales
: [SWD];
Lago Agrio:
[RWG].
Note: Often referred to as “near”
A. risi
,
then in the mid 1990’s it was referred to as “
A. dunklei
,” a manuscript name since it was thought to be a new species. Many specimens in various museums around the world are labeled “
A. dunklei
ms.” Another manuscript name that was proposed more recently was “
A. upano
;” Rosser Garrison (pers. comm., 2017) has determined that most of these refer to
A. limitata
,
a very common, mid-elevation east slope species. A few specimens labeled “
A. upano
ms.” turned out to be
A. philipi
Garrison & von Ellenrieder, 2018
.
Argia loutoni
Garrison & von Ellenrieder, 2015
. Publ. Recs.
ECU:
(P16),
Garrison & von Ellenrieder (2018)
;
SU, Shushufindi
: *
Garrison & von Ellenrieder (2015)
. Also BRA, COL, PER, VEN:
Garrison & von Ellenrieder (2015)
.
Addl. Data.
SU,
Shushufindi
:
1 ♀
, small stream about
3 km
W of Shushufindi
(
-0.1647
,
-76.6878
,
286m
),
28 x 2013
, leg. KJT, det. RWG [KJT].
Argia mauffrayi
Garrison & Ramón Cabrera, 2019
. Publ. Recs.
ECU: PC, San Miguel de Los Bancos Cantón:
Garrison & Ramón Cabrera (2019
—
Holotype
♂
, Recinto Milpe, Milpe Bird Sanctuary, Mirador Uno Trail (
0.0333
,
-78.8661
,
1101m
),
4 ix 2018
, leg. WFM [USFQ].
ES, Quinindé:
Paratype
♂
, Reserva Canande (
0.5203
,
-79.0866
,
470m
),
15 v 2012
, leg. Jessica Ware & Melissa Sanchez [RWG]).
Note: Known only from these two localities at present.
Argia medullaris
Hagen
in
Selys, 1865
. Publ. Recs.
ECU:
(H08), (P11), (H14);
IM:
Garrison & von Ellenrieder (2018)
;
LJ:
Garrison & von Ellenrieder (2018)
;
MN:
Garrison & von Ellenrieder (2018)
;
MS:
Garrison & von Ellenrieder (2018)
;
NA:
*
Tennessen (1996b)
, Daigle (1996),
Garrison & von Ellenrieder (2018)
;
PA:
Garrison & von Ellenrieder (2018)
;
PC:
Garrison & von Ellenrieder (2018)
;
SD:
Garrison & von Ellenrieder (2018)
;
TU:
Garrison & von Ellenrieder (2018)
. Also CEN, COL, VEN: (H08), (P11),
Garrison & von Ellenrieder (2018)
.
Addl. Data.
CT, Pangua:
[FSCA, KJT];
Sigchos:
[FSCA];
IM, Cotacachi:
[FSCA, KJT, USFQ];
San Miguel Urcuq:
[JTJ];
LJ, Catamayo:
[JTJ];
MS, Morona:
[FSCA, JJD, RWG];
NA,Archidona
: [FSCA, JTJ, KJT, SWD];
Quijos:
[FAL, FSCA, JJD, KJT, RWG, TWD, USNM];
PC, Mejía:
[FSCA, JJD, USNM];
San Miguel Bancos:
[FSCA, USNM];
SD:
[FAL, KJT];
SU, Gonzalo Pizarro:
[FSCA];
TU, Baños:
[RWG];
ZC, Zamora:
[JTJ].
Argia nigrior
Calvert, 1909
. Publ. Recs. ECU: (H08), (P11), (H14);
Garrison & von Ellenrieder (2018)
;
ZC:
*
Tennessen & Johnson (2008)
. Also PER: (H08), (P11).
Note: Rosser Garrison (pers. comm., 2017) wrote “A pair labeled
Argia
sp. “D”:
ECUADOR
:
Zamora-Chinchipe Province
,
5.6 km
east of Zamora,
3.7 km
south of Timbara [“small river with boulders, tributary of Río Zamora, end of gravel road, trickle on roadside and along trail bordering river, secondary forest and pasture”],
4.0658
,
-78.9044
;
4 iv 2008
; collected by Dave Wagner, William A. Haber, Michael Thomas, Fred Morrison & Ronald Vargas matches well the
type
of
A. nigrior
from Bolivia and have so labeled it this species.”
Argia oculata
Hagen
in
Selys, 1865
. Publ. Recs.
ECU:
*Calvert (1902),
Calvert (1902b)
,
Ris (1918)
,
Soukop (1954)
,
Paulson (1983)
,
Tsuda (1991)
,
Steinmann (1997a)
, (L06), (H08), (P11), (H14);
CR
:
Campos (1922)
;
CB
:
Calvert (1907)
,
Campos (1922)
;
EO
:
Campos (1927)
;
LR
:
Campos (1922)
;
AZ, BO, CT, ES, LJ, LR, PC, SD
:
Garrison & von Ellenrieder (2015)
,
MS
: Garrison & von Ellenrieder (2007);
NA:
Daigle (1996), Garrison & von Ellenrieder (2007);
OR
: Tennessen (1996), Garrison & von Ellenrieder (2007, 2011);
PA
: Garrison & von Ellenrieder (2007);
SU:
Garrison & von Ellenrieder (2007). Also BRA, COL, PER—as
A. difficilis
, TTO as
A. difficilis
, VEN: (L06), (H08), (P11); plus CEN, PER, TTO: (H08); and BOL, GUF, GUY, SUR: (P16).
Addl. Data.
AZ, Santa Isabel:
[FSCA, KJT, RWG, SWD];
BO, Caluma:
[FSCA, KJT, JTJ];
CB, Camada:
[RWG];
CN,
Cañar
:
[FSCA];
La Troncal:
[JJD, KJT, SWD];
San Miguel:
[FSCA, KJT];
CT, Pangua:
[FSCA, JTJ, KJT];
Sigchos:
[FSCA];
EO, Arenillas:
[FSCA];
Piñas:
[FSCA, JTJ, KJT];
Santa Rosa:
[JTJ, KJT];
ES, San Lorenzo:
[FSCA, JJD, KJT, RWG, USNM];
GY, Naranjal:
[FSCA, JTJ, KJT];
IM, Ibarra
: [FSCA];
LJ, Catamayo:
[FSCA, JTJ, KJT];
Loja
:
[FSCA, JTJ, KJT, UTIC];
LR, Buena Fé:
[SWD];
Quevedo:
[JTJ, KJT];
Montalvo:
[KJT];
MN, Flavio Alfaro:
[KJT];
Pedernales:
[JTJ, KJT];
PC, Mejía:
[FSCA];
San Miguel Bancos:
[FSCA, KJT, RWG, USFQ];
SD:
[FSCA, JJD, JTJ, KJT, RWG, SWD, USNM].
Note: Many specimens of
A. oculata
in various collections are labeled “
Argia
cf.
oculata
,” “
A. oculata
?,” “
A. difficilis
,” “
A. forficula
” and “
A. elipitca
”.
Garrison & von Ellenrieder (2015)
defined
A. oculata
and discussed its varability. It is very common in
Ecuador
on the Pacific slope.
Argia philipi
Garrison & von Ellenrieder, 2018
. Publ. Recs.
MS:
*
Garrison & von Ellenrieder (2018)
;
NA:
*
Garrison & von Ellenrieder (2018)
;
OR:
*
Garrison & von Ellenrieder (2018)
;
SU:
*
Garrison & von Ellenrieder (2018)
. Also BOL, COL, PER:
Garrison & von Ellenrieder (2018)
.
Addl. Data.
OR,
Loreto
:
2 ♂♂
, 1 cop. pair, forest stream about
12 km
NE of
Loreto
(
-0.6217
,
-77.2600
,
365m
),
12 x 2009
, leg. KJT [KJT]; same data but
1 ♂
, small streams and forest trails
12 km
E of
Loreto
, 2+ km
S of Hwy.
45, leg. JTJ, det. RWG [JTJ];
PA,
Mera
:
1 ♀
, near
Shell
, trails, forest, and
Río Metolo
, at
Los Copales
(
-1.4919
,
- 78.0725
,
1070m
),
20 ix 2005
, leg. KJT, det. RWG [KJT];
SU,
Shushufindi
:
1 ♂
, stream about
3 km
NW of Shushu- findi (
-0.1664
,
-76.6917
,
290m
),
13 x 2009
, leg. KJT, det. RWG [KJT].
Argia popoluca
Calvert, 1902
. Publ. Recs. ECU: (H14), *
Garrison & von Ellenrieder (2018)
. Also COL (as
A. variata
)
: (H08), (P11), (P&P).
Addl. Data.
BO, Caluma:
1 ♂
, tributary of rocky stream
5 km
E of Caluma, Gauranda Road (
-1.5942
-79.2308
,
480m
),
5 Mar 2010
, leg. JJD, det. RWG [JJD];
ES,
Quinindé
:
1 ♂
,
Río Canande Reserve
,
35 km
NE of Quininde
, tributary of
Río Hoja Blanca
, small gravel stream in forest (
0.5264
,
-79.2122
,
345m
),
20 x 2012
, leg. WAH, det. RWG, data provided by RWG (2018) [WAH];
PC,
Quito
:
2 ♂♂
,
2 ♀♀
,
Río Tulambi
,
Forestry Station
Maquipucuna (about
5 km
E of Nanegalito
) (
-0.1169
,
-78.6297
,
1344m
),
5 ix 1990
, leg. OSF, det. RWG [RWG, USNM];
1 ♂
,
2 km
NE of Mindo
,
Septimo Paraiso Lodge
, small stream in forest (
0.0308
,
-78.7633
,
1555m
), leg. WAH, det. RWG, data provided by RWG (2018) [WAH].
Argia pulla
Hagen
in
Selys, 1865
. Publ. Recs.
ECU:
(L06),
von Ellenrieder & Muzón (2007)
, (H08), (P11), (H14),
Garrison & von Ellenrieder (2018)
;
NA
: *Daigle (1996). Also BRA, CEN, COL, PER, TTO, VEN: (L06), (H08), (P11); plus ARG: (H08); and SUR: (P16).
Addl. Data.
ES,
Esmeraldas
:
[DRP];
MN, Chone:
[JJD, JTJ];
Flavio Alfaro:
[JTJ];
Junin
:
[JTJ, KJT];
MS,
Santiago
:
[SWD];
NA, El
Chaco
:
[SWD];
Tena:
[FSCA, JTJ, KJT, SWD, TWD, USNM];
OR, Francisco de
Orellana
: [FSCA, JJD, KJT, RWG, TWD, SWD, USNM];
Loreto
:
[JTJ];
SD:
[FSCA, JJD, KJT, USNM];
SU, Lago Agrio:
[SWD, USNM];
Shushufindi:
[FSCA, KJT, SWD, USNM].
Argia schneideri
Garrison & von Ellenrieder, 2017
. Publ. Recs.
MS:
*
Garrison & von Ellenrieder (2017)
;
NA:
*
Garrison & von Ellenrieder (2017
—
Holotype
: Las Palmas, on Río Anzu in Río
Napo
watershed,
11 xii 1936
, leg. WCM, [UMMZ];
PA:
*
Garrison & von Ellenrieder (2017)
;
TU:
*
Garrison & von Ellenrieder (2017)
. Also COL: Bota-Sierra
et al.
(2018).
Addl. Data.
OR
,
Loreto
: 1 cop. pair, forest stream about
12 km
NE of
Loreto
(
-0.6217
,
-77.2600
,
365m
),
12 x 2009
, leg. & det. KJT [KJT];
1 ♂
, same data but
15 x 2009
, leg. & det. JTJ [JTJ];
1 ♂
, small streams and forest trails
12 km
E of
Loreto
, 2+ km S of
Hwy.
45 (
-0.6217
,
-77.2600
,
365m
),
15 x 2009
, leg. & det. JTJ [JTJ];
SU
,
Shushufindi
:
1 ♂
, trails, small streams
3 km
W of Shushufindi
, 2+ km N off Hwy. 26 at km marker 17 (
-0.1664
,
-76.6917
,
290 m
),
13 x 2009
, leg. & det. JTJ [JTJ].
Note: The manuscript name “
camura
” was originally applied to this species.
Argia selsyi
Garrison & von Ellenrieder, 2018
. Publ. Recs.
MS:
*
Garrison & von Ellenrieder (2018)
;
NA:
*
Garrison & von Ellenrieder (2018
—
Holotype
: [
Tena]:
Jatun Yacu,
Napo
River watershed [
-1.0166
,
-77.8333
,
700m
],
18 iv 1935
, leg. WCM [UMMZ]);
OR:
*
Garrison & von Ellenrieder (2018)
;
PA:
*
Garrison & von Ellenrieder (2018)
;
TU:
*
Garrison & von Ellenrieder (2018)
.
Note: Endemic but common when found.
Argia talamanca?
Calvert, 1907
. Publ. Recs.
ECU:
*
Tennessen (1998)
, (P11), (H14). Also CEN, COL, VEN: (H08), (P11).
Addl. Data.
BO, San Miguel:
1 ♂
, Balzapamba (-1.77, -
79.18W
,
732m
),
ix 1935
, leg. R. Campos, det. L. Gloyd [FSCA];
CN, La Troncal:
2 ♂♂
, small stream and waterfall,
25 km
E of Cochancay, Hwy. 70, (
-2.4692
,
-79.1697
,
1540m
),
3 xi 1997
, leg. & det. JJD [FSCA];
4 ♂♂
, same data but leg. & det. KJT [FSCA];
5 ♂♂
,
2 ♀♀
, same data but leg. & det. TWD [RWG, TWD, FSCA];
3 ♂♂
, same data but leg. & det. WFM [FSCA, LSAM, USNM];
CT, Sigchos:
3 ♂♂
, San Francisco de las Pampas Otonga Reserve (-0.42, -79.0,
1700m
),
14–16 ii 1997
, leg. & det. Elicio Tapia, [FSCA, QZAC];
ES, San Lorenzo:
1 ♂
,
5 km
NW of Lita, seeps along RR tracks,
5 ii 1997
, leg. & det. JJD [FSCA];
MN, Jipijapa:
3 ♂♂
,
3 ♀♀
, Puerto Lopez “
20 km
to Salinas” (-1.57, -80.82,
61m
),
5 iv 97
, leg. & det. Elicio Tapia [FSCA, QZAC];
PC, Mejía:
1 ♂
, Manuel Cornejo
2 km
S of Tandapi, waterfalls (-0.43, -78.78,
1737m
),
21 x 1980
, leg. & det. MJW [FSCA];
1 ♂
, Manual Cornejo E near Tandapi, “east of Santo Domingo de los Colorados” (
-0.4167
,
-78.7833
,
1585m
),
21 x 1980
, leg. & det. MJW [FSCA];
7 ♂♂
, same data but
24 x 1980
. [FSCA];
TU, Baños:
1 ♂
, La Ventana, upper Río
Pastaza
watershed,
ix 1935
, leg. & det. F Campos R, [FSCA].
Note: South American specimens may represent a new undescribed species, since the true
A. talamanca
may occur only in Central America (R. Garrison, pers. comm., 2018). Very similar to
A variegata
.
Argia talamanca
appears to be a west slope species whereas
A. variegata
is mainly an east slope species.
Argia tennesseni
Garrison & von Ellenrieder, 2018
. Publ. Recs. NA: *
Garrison & von Ellenrieder (2018)
;
OR:
*
Garrison & von Ellenrieder (2018
—
Holotype
: [
Loreto
]: stream
8.5 km
E of
Loreto
[
-0.6183
,
-77.2950
,
400m
],
14 xi 1997
, leg. KJT [FSCA];
♀
Paratype
: stream on road to Waorani Reserve,
6.2 km
south of main Yasuni Road [
- 0.7120
,
-76.4744
,
230m
],
10 vi 1995
, leg. KJT [FSCA];
SU:
*
Garrison & von Ellenrieder (2018)
. Also BRA, PER:
Garrison & von Ellenrieder (2018)
.
Addl. Data.
OR,
Francisco
de
Orellana
:
3 ♂♂
,
1 ♀
, stream, road to
Waorani Reserve
,
6.2 km
S of Yasuni Sci. Res. Sta.
,
10 vi 1995
, leg. KJT [KJT, RWG];
1 ♀
, P.N.Y., on road to
Waorani Reserve
,
1.8 km
W of Waorani
Re- serve
Road
, on
Markus Oil Co. Road
PC 29-30, small stream (
-0.7000
,
-76.4667
,
200m
), leg. WFM, det. RWG & NVE [RWG];
1 ♂
,
2 ♀♀
, T.
B.S.
(
-0.6333
,
-76.1500
,
220m
),
20 ix 2000
, leg.
Kelly Jackson
&
D. Inward
, det. RWG [BMNH];
1 ♀
, T.
B.S.
, USFQ, stream crossing
Matapalo
trail at marker 450 (
-0.6369
,
-76.1458
,
245m
),
14 i 2009
, leg. & det. RWG & NVE [RWG];
1 ♂
,
4 ♀♀
, T.
B.S.
, USFQ, stream crossing
Chorongo
trail at marker 225 (
-0.6350
,
-76.1503
,
220m
),
16 i 2009
, leg. & det. RWG & NVE [RWG];
1 ♂
,
4 ♀♀
, same data but
18 i 2009
[RWG];
2 ♀♀
, T.
B.S.
, USFQ, black water stream crossing
Guacamayo
trail at marker 2050 (
-0.6383
,
-76.1492
,
220m
),
17 i 2009
, leg. & det. RWG & NVE [RWG];
2 ♀♀
, T.
B.S.
, USFQ,
Guacamayo
trail between markers 450–2100 (
-0.6350
,
- 76.1533
,
237m
),
22 i 2009
, leg. & det. RWG & NVE [RWG];
Loreto
:
3 ♂♂
, small streams and forest trails
12 km
E of
Loreto
, 2+ km S of
Hwy.
45 (
-0.6271
,
-77.2600
,
365m
),
12 x 2009
, leg. JTJ [JTJ];
1 ♂
,
2 ♀♀
, 1 reared, same data but leg. KJT [KJT].
Argia translata
Hagen
in
Selys, 1865
. Publ. Recs.
ECU
: (H08), (P11).
CR, GY, LR
: *
Campos (1922)
,
EO:
Campos (1927)
. Also ARG, CEN, COL, GUF, PER, TTO, VEN: (H08), (P11); plus BOL, GUY, SUR: (P16).
Addl. Data.
EO, Piñas:
[FSCA, JTJ];
Santa Rosa:
[KJT];
GU, Balao:
[FSCA, KJT];
Naranjal:
[FSCA];
LR, Buena Fé:
[DRP, FSCA, RWG, SWD];
Quevedo:
[FSCA, KJT];
Vantanas:
[FSCA, KJT];
MN, Chone:
[USNM];
Manta:
[RWG, USNM];
NA, Tena:
[FSCA, JJD, RWG, SWD, TWD, USNM]; SD: [RWG];
SU, Gonzalo Pizarro:
[USNM].
Argia variegata
Förster, 1914
. Publ. Recs.
ECU:
*
Förster (1914
- Description, no specific
type
locality),
Davies & Tobin (1984)
,
Tsuda (1986
,
1991
),
Bridges (1993)
,
Steinmann (1997a)
,
Garrison
et al.
(2003)
, (H08), (P11), (H14);
MS
:
Tennessen (2006)
;
NA
: Daigle (1996),
Tennessen (1998)
;
ZC
:
Tennessen & Johnson (2008)
. Also COL, PER: (H08), (P11).
Addl. Data.
MS, Huamboya:
[FAL, FSCA, JJD, KJT, TWD, USNM];
Morona:
[FSCA];
NA, Archidona:
[FAL, FSCA, JJD, JTJ, KJT, SWD, TWD, USNM];
Carlos Julio Arsemena Tola:
[RWG];
El
Chaco
:
[SWD];
Quijos:
[JJD];
Tena:
[SWD];
LJ,
Loja
[
JTA
]; OR,
Loreto
:
[DRP, FSCA, JJD, KJT, RWG, TWD, USNM];
PA, Mera:
[FSCA, JJD, KJT, RWG, UMMZ, USNM];
PC, San Miguel de Los Bancos:
[FSCA, USFQ];
SU, Gonzalo Pizarro:
[FSCA];
TU, Baños:
[CSCA, FSCA, RWG, UMMZ, USNM];
ZC, Zamora:
[FSCA, JTJ, KJT, USNM].
Note: This species is closely related to
A. talamanca
and may be found to be a junior synonym of it; the issue is currently under review by Rosser Garrison.
Argia
unresolved:
There are numerous specimens and records for
Argia
which we have not listed here. Many belong to the
Argia
gerhardi-kokama-nigrior
group that still needs to be resolved. Museum specimens are often labeled by alphabetical designation (e.g. “
Argia
sp. g”). Garrison & von Ellenrieder have been working on this complex for some time and hopefully someday will provide a solution for these unresolved
Argia
.