A Catalogue and Historical Study of the Odonata of Ecuador Author Mauffray, William F. Author Tennessen, Kenneth J. text Zootaxa 2019 2019-07-03 4628 1 1 265 journal article 26210 10.11646/zootaxa.4628.1.1 3e77e5cf-7e8a-490d-a5f7-c571b90aea57 1175-5326 3335707 E4C2F52F-9A1A-4C55-B03B-C87B2B4C4676 Libellula Linnaeus 1758 Of the 28 described New World species, only one species has been found in Ecuador ( Fig. 99 ), the nymphs of which develop in small shaded pools and ditches. Libellula herculea Karsch, 1889 . Publ. Recs. ECU: Kennedy (1922) , Racenis (1954) , Paulson (1983) , Tsuda (1986 , 1991 ), (H06), (P11), (H14); AZ : * Kirby (1889 —as Belonia longipennis ), Calvert (1906) , Campos (1922) ; BO : Ris (1910) ; ES: Calvert (1909) , Campos (1922) , Daigle et al. (1997) ; OR : von Ellenrieder & Garrison (2011) ; PC: * Calvert (1909) , Campos (1922) . Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, VEN: (H06), (P11); plus SUR: (P16). FIGURE 99. Libellula herculea ♂, ECUADOR, Orellana Province, ditch-side stream S of Cascada de Pasahurco, 16.6 km W of Loreto, 4 xi 2013; photo K. J. Tennessen. Addl. Data. AZ , Santa Isabel: 1 ♂ , stream, 21 km E of Hwy. 25, road from Balao jct. ( -2.9400 , -79.5653 , 730m ), 11 iv 2008 , leg. & det. KJT [KJT]; 1 ♂ , 1 ♀ , same data but leg. & det. JTJ; BO, San Miguel: 1 ♂ , Bal- zapamba ( -1.7667 , -79.1833 , 700m ), ix 1935 , leg. F. Campos , det. MJW [ANSP]; EO, Piñas : Buenaventura , Río Principal ( -3.6561 , -79.7735 , 329m ), 19 vi 2016 , leg. & det. ACRP [ACRP]; ES , San Lorenzo : Lita along RR track, 1–5 km W, adjacent to Río Baboso seeps, pools & streams ( 0.8792 , -78.4589 , 535m ), 5 ii 1997 , 5 ♂♂ , 1 ♀ , leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 1 ♂ , 1 ♀ , same data but leg. & det. JJD [JJD]; 1 ♂ , same data but leg. & det. KJT [KJT]; MS, Huanboya : 1 ♂ , small stream near Río Pasanac , Hwy . 45, S of river, E side of Hwy . , 16 ix 2005 , leg. & det. KJT [KJT]; 1 ♀ , Macas , 58 km N on Hwy . 45 ( 70 km N of Sucua ), roadside ditch and tribs near Río Pasanac ( -1.9447 , -77.8536 , 762m ), 10 xi 1997 , leg. & det. WFM [FSCA]; Taisha : 1 ♀ , Macuma , small ponds ( 867m ), 1 ii 1972 , leg. D. Pearson , det. DRP [DRP]; NA , Tena : 2 ♂♂ , Tena ( 500m ), leg. F. Campos , det. MJW [ANSP]; 1 ♂ , Tena (-0.98, -77.80, 500m ), date?, leg. F. Campos , det. MJW [FSCA]; adult, 1 km S of Tena , 28 v 1977 , leg. P. Spangler & D. Givens , det.? [USNM]; 1 ♂ , J. S.B. S., near Tena ( 450m ), 18 vi 1994 , leg. A. Jaramillo , det. DRP [DRP]; OR , Francisco de Orellana : 1 ♂ , T. B.S. , open grassy area behind dining room, surrounded by bushes and trees and some seepage areas ( -0.6383 , -76.1492 , 220m ), 10 i 2009 , leg. & det. RWG & NVE [USFQ]; Loreto : 1 ♀ , small streams and forest trails 12 km E of Loreto , 2+ km S of Hwy. 45 ( -0.6217 , -77.2600 , 365m ), 12 x 2009 , leg. & det. JTJ [JTJ]; 1 ♂ , Cotapino, S along roadside stream ( -0.7076 , -77.4743 , 530m ), 4 xi 2013 , leg. & det. CDK [FSCA]; same data but 1 ♂ , leg. & det. KJT [FSCA]; PC , San Miguel Bancos : 1 ♂ , Puerto Quito ( 750m ), 28 v , leg.?, det. SWD [QZAC]; 1 ♂ , 1 ♀ , Mindo, swamps and streams near town ( -0.0556 , -78.7714 , 1244m ), 28 i 2009 , leg. & det. RWG & NVE [RWG]; 1 ♂ , 1 ♀ , Mindo, Via al Mariposario, Mindo Gardens Eco Lodge, vege- tated cement pond ( -0.0720 , -78.7560 , 1390m ), 3 ix 2018 , leg. & det. KJT [KJT]; Quito : 3 ♂♂ , 1 ♀ , Río Tulambi , Forestry Sta. Maquipucuna , about 5 km E of Nanegal (0.12, -78.62), 9 v 1990 , leg. OSF, det.? [USNM]; SD : 1 ♀ , Santo Domingo , 12 km E at Tinalandia , 11–17 v 1986 , leg. G. Edwards , det. SWD [SWD]; 2 ♂♂ , about 93 km SW Quito , Chiriboga Road , 9 vi 1990 , leg. OSF, det.? [USNM]; SU , Shushufindi : 1 ♀ , Limoncocha , forest trail, 8 x 1971 , leg. D. Pearson , det. DRP [DRP]. Note: There is a specimen in QZAC labeled “Calvla” 12 i 1977 ; we could not find this location in any reference. Macrothemis Hagen, 1868 A species-rich genus of 41 species with at least 14 species in Ecuador . The latest key to males was provided by May (1998) although species identification is difficult; females of very few species are identifiable. These are small to medium-sized mostly dark dragonflies ( Fig. 100 ) that inhabit small streams to medium-sized rivers, both open and forested. Macrothemis delia Ris, 1913 . Publ. Recs. E CU: *(H14). Also CEN, SUR, VEN: (H06), (P16). Addl. Data. OR , Francisco de Orellana : 1 ♂ , Río Shiripuno , bridge crossing 59 km SSE of Coca , ( -1.0079 , -76.9451 ), date not provided, data provided by WAH & DLW; 1 ♂ , T. B.S. , Río Tiputini , 55 km SE of Pompeya , 94 km ESE of Coca ( -0.6369 , -76.1503 ), date not provided, data provided by WAH & DLW. * Macrothemis extensa Ris, 1913 . Publ. Recs. BRA, CEN, GUF, PER, VEN: (H06), (P11); plus SUR: (P16). Addl. Data. LR , Buena Fé : 1 ♂ , Río Palenque Biological Station, 47 km S of Santo Domingo de los Colorados ( 0.5864 , -79.3647 , 150m ), 8 x 1988 , leg. & det. SWD [SWD]; SD: 1 ♀ , Alluriquín 7.3 km W of Hotel Tinalandia, 29 x 1994 , leg. A. Jaramillo, det. DRP [DRP]. * Macrothemis fallax May, 1998 . Publ. Recs. CEN: May (1998) , (P16); plus COL: (P&P), (P16). Addl. Data. ES , San Lorenzo : 1 ♀ , tiny stream near RR track, about 4 km N of Lita, 5 ii 1997 , leg. & det. KJT [KJT]; MN, Portoviejo : 1 ♂ , concrete ditch, Hwy. E 40, E of Portoviejo , 4 iii 2010 , leg. & det. JJD [JJD]; 1 ♂ , same data but leg. KJT, det. WFM [FSCA]; SD: 3 ♂♂ , Tinalandia , 12 km E of Santo Domingo de los Colorados , streams S of river (-0.2839, -79.05, 760m ), 5 x 1988 , leg. & det. SWD [RWG, SWD]. Note: The additional records are the southernmost for this species which is often confused with M. pseudimitans . FIGURE 100. Macrothemis hahneli ♂, ECUADOR, Napo Prov., small stream <1 km E of Río Sardinas Grande, 24 x 2013; photo K. J. Tennessen. Macrothemis hahneli Ris, 1913 . Publ. Recs. ECU: (H06), (P11), (H14). * Daigle et al. (1996) . NA : Daigle (1996— misidentified as M. musiva ). Also in COL, PER, VEN: (H06), (P11); plus ARG: (Garrison pers. comm. 2019). Addl. Data. MS , Huamboya : 1 ♂ , Macas, 58 km N on Hwy. 45 [ 70 km N of Sucua], roadside ditch and tribs near Río Pasanac ( -1.9447 , -77.8536 , 762m ), 16 ix 2005 , leg. & det. WFM [FSCA]; Morona : 3 ♂♂ , 1 ♀ , small stream and roadside seepage pool, 15 km W of Macas ( -2.2731 , -78.1939 , 1070m ), 15 ix 2005 , leg. & det. KJT [KJT]; 3 ♂♂ , 1 ♀ , leg. & det. WFM [FSCA, QZAC, USNM]; 3 ♂♂ , rocky stream, 22.5 km W of Macas, Guamote Road, 18 ix 2005 , leg. & det. JJD [JJD]; 2 ♂♂ , 2 ♀♀ , Macas, 22.9 km W on Guamote Road, 3.3 km W on Río Al- banico rocky stream and flowing roadside ditches ( -2.2342 , -78.2181 , 1690m ), 18 xi 2005 , leg. & det. WFM [FSCA, LSAM]; NA , Archidona : 5 ♂♂ , 1 ♀ , [Narupa], E on road to Loreto , 43.5 km W of Loreto seep near large rocky river [Río Hollin] ( -0.7022 , -77.7189 , 1036m ), 15 vii 1996 , leg, & det. JJD [JJD]; 2 ♂♂ , same data but leg. & det. SWD [SWD]; 1 ♂ , same data but 14 vi 1995 , leg. & det. KJT [KJT]; 1 ♂ , 1 ♀ , same data but 15 vii 1996 , leg. & det. WFM [FSCA, LSAM]; 5 ♂♂ , same data but 14 xi 1997 [FSCA, LSAM, QZAC, USNM]; 2 ♂♂ , [Sarayacu], 12.8 km N of Narupa on Baeza Hwy., small shallow sandy stream (-0.67, -77.78, 1585m ), 14 vii 1996 , leg. & det. WFM [LSAM, QZAC]; 3 ♂♂ , same data but leg. KJT [KJT]; 3 ♂♂ , same data but leg. & det. SWD [SWD]; 2 ♂♂ , seep by waterfall, Loreto Road, 44.1 km E of Narupa jct., 14 xi 1997 , leg. & det. KJT [KJT]; 1 male same data but leg. & det. TWD [SWD]; 2 ♂♂ , [Jondachi], 42 km N of Tena on road between Tena & Baeza (-0.72, -77.78, 1190m ), 27 x 1980 , leg. & det. MJW [FSCA, RWG]; Quijos : 1 ♂ , 1 ♀ , Baeza 4.8–5.0 km S, small clear streams and seeps ( -0.4761 , -77.8711 , 1830m ), 20 xi 1997 , leg. & det. WFM [FSCA]; 2 ♂♂ , same data but leg. & det. JJD [JJD]; 5 ♂♂ , 2 ♀♀ , same data but leg. & det. KJT [KJT]; 3 ♂♂ , 2 ♀♀ , same data but leg. & det. TWD [SWD]; 1 ♀ , same data but 10 v 2009 , [KJT]; 1 ♂ , same data but 3 xi 2001 , leg. & det. F. Sibley [FSCA]; 1 ♂ , Baeza, 1 mile S on Hwy. 45 ( -0.4641 , -77.8842 , 1830m ), 3 xi 2001 , leg. & det. F. Sibley [FSCA]; 1 ♂ , stream, Baeza-Tena Road, near San Isidore Road, 24 vii 1996 , leg. & det. KJT [KJT]; PC, Mejía: 1 ♀ , Tatatambo, about 5 km E on Hwy. 30, rocky stream (-0.45, -78.73, 2286m ), 30 i 1997 , leg. & det. WFM [FSCA]; PA , Mera : 1 ♀ , seep near Río Anzu, 12.0 km N of Mera ( -1.4181 , -78.0528 , 1190m ), 21 ix 2005 , leg. & det. KJT [KJT]; 1 ♂ , [Mera] Baños-Puyo Road, [about 5 km W], small forested pond ( -1.4806 , -78.1444 , 1097m ), 8 xi 2001 , leg. & det. F. Sibley [FSCA]; ZC, Yantzaza : 1 ♀ , Yantzaza, 10.8 km N [on Hwy. 45], rocky gravel stream ( -3.7669 , -78.6915 , 762m ), 6 xi 1997 , leg. & det. WFM [FSCA]; Zamora : 1 ♂ , stream/waterfall W of Zamora ( -3.9583 , -79.0289 , 1660m ), 1 iv 2008 , JTJ [JTJ]; 1 ♀ , 30 km E of Loja , foot of cascade, 23 ix 1990 , leg. OSF, det.? [USNM]. Macrothemis hemichlora ( Burmeister, 1839 ) . Publ. Recs. ECU: * Calvert (1906) , Ris (1913a) , Campos (1922) , Paulson (1983) , Tsuda (1986 , 1991 ), (H06), (P11), (H14). Also ARG CEN, COL, GUF, PER, SUR, TTO, VEN: (P11); plus BRA: (H06). Addl. Data. EO , Arenillas: [JTJ, KJT]; Piñas: [ACRP, FSCA]; Portovelo: [FSCA]; LR , Buena Fé: [SWD]; MN, Flavio Alfario: [FSCA]; MS , Morona: [FSCA, KJT]; Santiago : [SWD]; NA , Tena: [DRP, FSCA, UTIC, KJT, LSAM, QZAC, SWD, TWD, USNM]; OR , Francisco de Orellana : [KJT]; SU , Shushufindi: [FSCA, KJT]; ZC : Nangaritza: [FSCA]; Yantzaza: [FSCA]. * Macrothemis idalia? Ris, 1919 . Publ. Recs. GUY, VEN: (H06), (P11). Addl. Data. OR , Loreto : 1 ♀ , 12 km NE of Loreto , forest ( -0.6217 , -77.2600 , 365m ), 12 x 2009 , leg. KJT, det. RWG [FSCA]; SU, Lago Agrio : 2 ♀♀ , [ Santa Cecillia ]. 24 km W of Lago Agrío on road to Baeza , stream, 20 viii 1980 , leg. SWD, det. WFM [FSCA]. Note: Rosser Garrison (pers. comm.) compared the from near Loreto with ♀♀ in his collection and tenta- tively identified it as this species. A male from Sucumbíos Province , near Shushufindi, collected by KJT on 13 May 2009 was sent to R. W. Garrison who replied “... this could be the unknown ? of M. idalia Ris – size, body pattern, claws etc. consistent with known females; but until new material, association would be difficult.” Further collecting, especially a mated pair, will be necessary to confirm this record. Heckman (2006) stated that “without a description of the , the status of this species is uncertain.” * Macrothemis imitans Karsch, 1890 . Publ. Recs. ECU: (H14). Also ARG, BOL, BRA, CEN, GUY, PRY, TTO, URY, VEN: (H06), (P11); plus COL: (P&P). Addl. Data. NA , Tena : 3 ♂♂ , 1 ♀ , Puerto Napo 5.3 km E of Hwy. 45 on J.S.B.S. Road, Río Sinde ( -1.0500 , - 77.7481 , 450m ), 15 vi 1995 , leg. & det. WFM [FSCA, QZAC, USNM]; 3 ♂♂ , same data but leg. & det. KJT [KJT]; 2 ♂♂ , same data but leg. & det. SWD [SWD]; 1 ♂ , same data but 13 vii 1996 [KJT]; 2 ♂♂ , 1 ♀ , same data but leg. & det. WFM [FSCA, QZAC]; 3 ♂♂ , Puerto Napo 11.2 km E of Hwy. 45 on J.S.B.S. Road, Río Tiyuyacu ( -1.0516 , -77.7017 , 396m ), 13 xi 1997 , leg. & det. WFM [FSCA, QZAC, USNM]; OR , Francisco de Orellana : 1 ♂ , [Pom- peya Sur] S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (-0.60, -76.47, 229m ), 17 xi 1997 , leg. & det. WFM [FSCA]; PA , Pastaza : 1 ♂ , Río Sandalias , 5 km S of Puyo , Hwy. 45 ( -1.5003 , -77.9522 , 1000m ), 11 xi 1997 , leg. & det. KJT [KJT]; 1 ♂ , same data but leg. & det. WFM [FSCA]; SU , Cascales : 1 ♂ , river, 37 km W of Lago Agrio , 20 viii 1980 , leg. & det. SWD [SWD]; Lago Agrio : 1 ♂ , stream, 26 km S of Lago Agrio , 19 viii 1980 , leg. & det. SWD [SWD]. Macrothemis inequiunguis Calvert, 1895 . Publ. Recs. ECU: * Calvert (1898 , 1906 ), Ris (1913a) , Campos (1922) , Tsuda (1991) , (H06), (H14). Also CEN, VEN: (H06), (P11); plus COL: (H06). Addl. Data. SD: 2 ♂♂ , (as M. cf. inequiunguis ), Alluriquín 7.3 km W, forest stream at Hotel Tinalandia, 20 vii 1977 , leg. & det. DRP [DRP]. Note: The distribution data indicate that M. inequiunguis is confined to the west slope, its range extending northward to Central America and extreme southern United States , whereas M. tessellata (see below), a very similar species considered by some to be synonymous with M. inequiunguis , is confined to the east slope. * Macrothemis ludia Belle, 1987 . Publ. Recs. ECU: (H14). Also VEN: (H06), (P11); plus SUR: (P16). Addl. Data. NA , Tena , 1 ♀ , J.S.B.S., near Tena, in garden ( 450m ), 12 vi 1994 , leg. A. Jaramillo, det. DRP [DRP]; OR , Francisco de Orellana : 1 ♂ , T. B.S. , Río Tiputini , 55 km SE of Pompeya , 94 km ESE of Coca ( -0.6369 , -76.1503 ), date not provided, data provided by WAH & DLW; SU, Shushufindi : 1 ♂ , Limoncocha , forest stream, 9 iii 1972 , leg. D. Pearson , det. DRP [DRP]. Macrothemis mortoni Ris, 1913 . Publ. Recs. ECU: (H06), (P11), (H14); NA : * Tennessen (1996b) . Also BOL PER, VEN: (H06), (P11). Addl. Data. NA , Archidona : 1 ♂ , stream 43.5 km W of Loreto ( 1100m ), 14 vi 1995 , leg. & det. KJT [KJT]; Tena : 1 ♂ , stream, road to J.S.B.S., 11.2 km E of Puerto Napo , 15 vi 1995 , leg. & det. KJT [KJT]; 1 ♂ , Puerto Napo , 3.2 km E on J.S.B.S. Road, sloughs along Napo River, 23 vii 1996 , leg. & det. SWD [SWD]; MS , Huamboya : 1 ♂ , stream 10 km S. of Comunidad Chuwitayo ( -1.9500 , -77.8583 , 800m ), 10 xi 1997 , leg. & det. TWD [SWD]. Macrothemis musiva Calvert, 1898 . Publ. Recs. ECU: Schmidt (1942) , (H06), (P11), (H14); ES : * Ris (1913a) , Campos (1922) . Also ARG, BOL, BRA, CEN, COL, GUF, PRY, PER, VEN: (H06), (P11). Addl. Data. MS , Gualaquiza : 3 ♂♂ , 3 ♀♀ , Gualaquiza (-3.40, -78.58, 800m ), iv–v 1935, leg. F. Campos, det. MJW [ANSP, FSCA]; NA , Tena : 4 ♂♂ , 1 ♀ , Puerto Napo 3.2 km E on J.S.B.S. Road ponds and flowing marsh ( -1.0456 , -77.7467 , 420m ), 23 vii 1996 , leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 2 ♂♂ , same data but leg. & det. KJT [KJT]; 2 ♂♂ , same data but leg. & det. SWD [SWD]; 1 ♀ , Puerto Napo 5.3 km E of Hwy. 45 on J.S.B.S. Road, Río Sinde ( -1.0500 , -77.7481 , 450 m ), 22 vii 1996 , leg. & det. WFM [FSCA]; 1 ♀ , Puerto Napo 27.7 km E of Hwy. 45 on J.S.B.S. Road, Río Chinchipino ( -1.0728 , -77.5864 , 385m ), 13 xi 1997 [FSCA]; 2 ♀♀ , same data but leg. & det. KJT [KJT]; 1 ♂ , Tena, 13 ix 1990 , leg. OSF, det.? [USNM]; OR , Francisco de Orellana : 1 ♂ , P.N.Y., 2.3 km E of Yasuni Biological Research Sta. , small stream (-0.67, -76.37), 9 vi 1995 , leg. & det. WFM [FSCA]; 1 ♂ , P.N.Y., 3.6 km S of Yasuni Road on Waorani Reserve Road ( -0.6956 , -76.4594 , 244m ), 10 vi 1995 , leg. & det. WFM [QZAC]; 1 ♂ , P.N.Y., Yasuni Biological Research Sta. along trails and roads within 5 km of Res. Sta. ( -0.6719 , -76.3992 , 244m ), 18 vii 1996 , leg. & det. WFM [FSCA]; 1 ♂ , stream, Yasuni Road , 20.5 km N Sci. Res. Sta. , 11 vi 1995 , KJT [KJT]; Loreto : 1 ♂ , pond & small stream 34 km W of Loreto ( 835m ), 14 vi 1995 , leg. & det. SWD [JJD] 4 ♂♂ , Pano , 12 ix 1990 , leg. OSF, det.? [USNM]; SU , Cascales : 2 ♂♂ , stream 58 km W of Lago Agrio , El Paraiso ( 455m ), 8 vi 1995 , leg. & det. KJT [KJT]; Lago Agrio : 2 ♂♂ , stream, E edge of Lago Agrio , 18 viii 1980 , leg. & det. SWD [SWD]; PA , Pastaza : 1.5 km S of Puyo , 19 v 1977 , leg. P. Spangler & D. Givens , det.? [USNM]. Macrothemis pseudimitans Calvert, 1898 . Publ. Recs. ECU: Muttkowski (1910) , Maria (1938) , Paulson (1983) , Tsuda (1986 , 1991 ), (H06), (P11), (H14); EO : Campos (1927) ; ES : Campos (1922) ; IM : Ris (1913a) , Campos (1922) ; GY : * Calvert (1906) , Campos (1922) . Also BRA, CEN, COL, TTO, VEN: (H06), (P11); plus PER: (P16). Addl. Data. IM, Ibarra: 3 ♂♂ , “Guayupe” [presumably “Guallupe”] (-0.78, -78.30, 1000m ), 13 ii 1946 , leg. L. Alonzo, det. MJW [FSCA]; 1 ♂ , Lita, 45.8 km E, rocky stream (-0.78, -78.32, 855m ), 6 ii 1997 , leg. & det. WFM [FSCA]. Note: It is possible that some of the previous records for this species are actually for the very similar M. fallax , which was not described until 1998. Macrothemis tessellata ( Burmeister, 1839 ) . Publ. Recs. ECU: * Paulson (1983) , Tsuda (1986 , 1991 ), (H06), (P11), (H14). Also ARG, BRA, CEN, COL, PRY: (P11); plus SUR, VEN: (P16). Addl. Data. NA , Tena : 1 ♂ , Río Sinde, 5 km E of Puerto Napo ( -1.0500 , -77.7481 , 450m ), 15 i 1995 , leg. & det. KJT [KJT]; OR , Francisco de Orellana : 2 ♂♂ , P.N.Y., 3.6 km S of Yasuni Road on Waorani Reserve Road ( -0.6956 , -76.4594 , 244m ), 10 vi 1995 , leg. & det. WFM [FSCA, QZAC]; 1 ♀ , P.N.Y., Yasuni Biological Research Sta. along trails and roads within 5 km of Res. Sta. ( -0.6719 , -76.3992 , 244m ), 18 vii 1996 , leg. & det. WFM [FSCA]; PA , Mera : 1 ♂ , Río Anzu 12.0 km N of Mera , 21 ix 2005 , leg. & det. JJD [JJD]; SU, Shushufindi : 1 ♂ , trails, small streams 3 km W of Shushufindi , 2+ km N off Hwy . 26 at km marker 17 ( -0.1664 , -76.6917 , 290m ), 14 x 2009 , leg. & det. JTJ [JTJ]; 1 ♂ , Limoncocha , 23 viii 1980 , leg. & det. SWD [SWD]; 1 ♂ , Shushufindi , about 3 km W, small stream, ( -0.1662 , -76.6879 ), 28 x 2013 , leg. & det. KJT [FSCA]. Note: We are treating these specimens from the east slope, previously determined to be M. inequiunguis , as M. tessellata ; see also note under M. inequiunguis . Macrothemis spp. unresolved: IM, Cotacachi: 1 ♀ , Santa Rosa , El Refugio de Intag Cloud Forest , south along road at roadside seeps ( 0.3693 , -78.4728 , 1854m ), 23 viii 2018 , leg. & det. WFM [FSCA]; same data but 1 ♂ , roadside seeps and open small open stream ( 0.3703 , -78.4745 , 1866m ), 23 viii 2018 [FSCA]; 2♂♂ , 2♀♀ , same data but leg. KJT [KJT]; PC, Quito : 1 ♂ , 1 ♀ , Mindo ( 1354m ), 15 v 2011 , leg. & det. ACRP [ACRP]; SU, Shushufindi : 1 ♀ , Shushufindi road, 2.5 km W of Shushufindi at km marker 18, 3.5 km S, ( -0.2136 , -76.6786 , 280 m ), 12 v 2009 , leg. KJT (specimen labeled M. newtoni ?); 1 ♂ , 1 ♀ , Shushufindi , road, about 3 km NW of Shushufindi , ( -0.1664 , -76.6167 , 290m ), 13 v 2009 , leg. KJT (both of these specimens were determined by RWG as M. cf. brevidens ). Miathyria Kirby, 1889 A genus of just two species, easily distinguished ( Ris 1913b ). Both species are found in lentic waters such as ponds and canals with floating aquatic plants, especially water hyacinth and water lettuce; the nymphs hide in the roots. See Fig. 101 . FIGURE 101. Miathyria marcella ♀, UNITED STATES, Alabama, Fort Morgan Historic Site, 25 xi 2017; photo K. Chias- son. Miathyria marcella (Selys in Sagra, 1857) . Publ. Recs. ECU: Maria (1938) , Paulson (1983) , Tsuda (1986 , 1991 ), (H06), (P11), (H14); CR : Campos (1922) ; CB : Campos (1922) ; EO : Campos (1927) ; GY : * Therese (1900) , Calvert (1906) , Ris (1913b) , Campos (1922 , 1930 , 1932 ); LR : Calvert (1906) , Campos (1922) . Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, TTO, URY, VEN: (H06), (P11); plus SUR: (P16). Addl. Data. BO, San Miguel: [USNM]; CN, Cañar : [FSCA]; EO, Pasaje: [QZAC]; GY, Duran: [ANSP]; Guayaquil: [ANSP, FSCA, USNM]; Naranjal: [FSCA]; LR , Babahoyo: [USNM]; Buena Fé: [DRP]; Ventanas: [JJD]; MN : Chone: [USNM]; Sucre : [USNM]; SU , Shushufindi: [FSCA]. Miathyria simplex ( Rambur, 1842 ) . Publ. Recs. ECU: Ris (1913b) , Paulson (1983) , Tsuda (1986 , 1991 ), (H06), (P11), (H14); EO : Campos (1927) ; GY : * Calvert (1906) , Campos (1922 , 1930 ). Also BRA, CEN, GUF, GUY, PER, VEN: (H06), (P11); plus TTO: (P11); and COL, SUR (P16). Addl. Data. GY, Guayaquil: 1 ♀ , Guayaquil, 18 viii 1917 , leg. F. Goding, det.? [USNM]; OR , Francisco de Orellana : 1 ♂ , [ La Nueva Esperanza ], [across Río Napo from], near Primavera “Limoncocha”, Río Napo & Lake Taracoa (-0.45, -76.77, 300m ), 26 vii 1980 , leg. & det. SWD [SWD]; 3 ♂♂ , P.N.Y., Río Savaleto 2 & slough at km 31, 17 vii 1996 , leg. & det. SWD [SWD]; 2 ♂♂ , trib. of Río Payamino , road to Loreto , 1.7 km W of jct. with Coca Road ( -0.4367 , -77.0017 , 258m ), 18 xi 1997 , leg. & det. KJT [KJT]; 2 ♂♂ , same data but leg. & det. WFM [FSCA, QZAC]; 1 ♀ , Puerto Francisco de Orellana , 11.8 km W on Loreto Road, large pond ( -0.4742 , -77.0583 , 285m ), 18 xi 1997 , leg. & det. WFM [FSCA]; 1 ♀ , same data but leg. & det. KJT [KJT]; SU , Lago Agrio : 1 ♀ , 2 km N of Lago Agrio , 27 viii 1975 , leg. A. Langley , det.? [USNM]; Shushufindi : 6 ♂♂ , 2 ♀♀ , Limoncocha , Río Napo ( -0.4000 , -76.6167 , 269m ), 6 xi 1980 , leg. & det. MJW [FSCA]; 2 ♀♀ , same data but 25 xi 1980 [FSCA]; 1 ♀ , same data but 3–16 vi 1977 , leg. P. Spangler & D. Givens , det.? [USNM]; 1 ♂ , Limoncocha , clearing, 24 vii 1977 , leg. & det. D. Paulson [DRP]; 2 ♂♂ , Limoncocha , lakeshore, 26 vii 1977 , leg. DRP & S. Hills , det. DRP [DRP]; 1 ♂ , same data but 25 viii 1980 , leg. & det. SWD [SWD].