Revision of the Oriental species of Calleida Latreille (sensu lato). Part 1: Introduction, groups of species, and species of six species groups (Coleoptera: Carabidae: Lebiini)
Author
Casale, Achille
Author
Shi, Hongliang
text
Zootaxa
2018
2018-06-29
4442
1
journal volume
29772
10.11646/zootaxa.4442.1.1
ded8bdb6-f8ba-4e0b-bed1-1ec08963edfd
1175-5326
1302713
195B0471-553A-4617-B901-E9DBD2323D14
[11]
Calleida rapax
Andrewes, 1933
Calleida rapax
Andrewes, 1933
: 399
(
Type
locality:
India
: Dehra
Dun
, Lachiwala;
holotype
deposited in NHML).
Type
material examined
:
Holotype
,
male
, “
Lachiwala, Dehra
Dun U.P.,
C.F.C. Beeson
.,
6.IV.1923
”,
“Tectona grandis”, "Under bark
", "326", "
India
.
Dehra Dun
Coll.
B.M 1932-26
", "
Calleida rapax Andr.
Type H.E. Andrewes det.", "Type" (NHML,
fig. 78
).
Paratypes
: one male, "Nauri, 2,500’ Landsdowne. B.M. Bathia.
18.III.1926
”, “For. Res. Inst. Dehra Duna”, “dry ex Bamboo”, "305", "Co-type", “Andrewes coll. B.M.
1945-97
”, “
Calleida rapax Andr. Cotype H.E Andrewes
det.” (
MHNL
); one female, "New Forest, Dehra
Dun
, U.P. R.N. Mathur.
6.IX.1932
", "1590", "Co-type", "For. Res. Inst. Dehra
Dun
", "
Calleida rapax Andr. Cotype, Andrewes
det.", "H.E. Andrewes Coll. B.M.
1945-97
." (
MHNL
).
Non-type material examined:
1 ex., “
India
, Dehra
Dun
. G. Rogers 1906-202” (
CCA
, ex S.L Straneo).
Taxonomic note.
This taxon is very similar to
C. doriae
. They only have small differences in colour and in the apex of the aedeagal median lobe. Considering their allopatric distribution,
C. rapax
could be treated as a mere northwestern isolated subspecies of
C. doriae
, or a synonym of it. However, owing to the scarce examined material, at the moment we treat it as a valid species.
Its relationships suggested by
Andrewes (1933)
are wrong: “very closely allied to
femoralis
Chaud.,
sultana
Bates
, and
pallipes
Andr.
”. This species was not cited by
Jedlička (1963)
in his monograph.
Diagnosis.
See above, in the key to the species of the
C. doriae
species group and diagnosis of
C. doriae
.
Description.
The original description provided by
Andrewes (1933)
, here completed with some additional characters, is enough to distinguish this species.
General features
: As in
fig. 78
. Large sized species: L: 12.0–13.0 mm.
Colour
: Piceous to reddish brown, shiny; head reddish brown, occiput darker; antennomere 1 yellowish brown, antennomeres 2, 3 and basal half of 4 reddish, apical half of antennomere 4 and lateral sides of remaining antennomeres dark; base and apex of palpomeres yellowish; pronotal disc dark brown, with sides widely reddish; elytra metallic green with faint bluish purple discal patch, sometimes the marginal furrows greenish blue or violet; major parts of legs blackish, trochanters reddish, fourth and fifth tarsomeres reddish brown; underside brownish or black.
Lustre and microsculpture:
microsculpture vanished on head and pronotum; faint, formed by isodiametric meshes on elytra.
Head
flat and impunctate; frontal foveae shallow and uneven; eyes prominent; neck constriction evident; antennae reaching backwards to the basal fifth of elytra.
Pronotum
rather flat and cordate, transverse (ratio PW/PL: 1.13-1.19), a little wider than the head; base oblique at sides, wider than anterior margin. Anterior angles rounded; lateral sides moderately explanate and slightly reflexed, rounded in front, then nearly straight to the posterior angles which are obtuse, not projecting laterally. Disc smooth; median furrow and basal foveae fairly deep, the foveae continued forward on each side forming shallow sulci, parallel with lateral sides.
Elytra
depressed, three-fifths wider than pronotum, nearly four-fifth longer than wide; lateral sides slightly compressed behind shoulders and then a little dilated, so that the widest point is behind middle. Elytral apex obliquely truncate, the outer apical angles markedly widened, obtusely rounded, not angulate. Striae moderately deep, very finely punctate; intervals moderately convex, with some sparse minute punctuation.
Underside
with very fine scattered punctuation and pubescence on metasternum, more developed in males. Abdominal sternum VII in males notched at middle, with one seta on each side, two setae in females.
Male genitalia
very similar to that of
C. doriae
, but apex of the median lobe of aedeagus a little shorter (
fig. 77
); median lobe slightly bent, its middle part strongly expanded in dorsal view; dorsal and ventral margins slightly curved in lateral view, ventral surface markedly concaved in the middle; apical orifice pleuropic left, with a triangular chitinized piece; apical lamina flat, rounded at apex, expanded and thickened, basal width greater than length (W/L about 1.4); endophallus with two chitinized copulatory pieces, located in the middle area near the left lateral margin, long and narrow, close to each other at base, V shaped; left paramere depressed on the dorsal side; right paramere not curved at apex.
Female genitalia
: not examined.
Geographical Distribution and habitat.
Only known from northwestern
India
(
map 6
):
Uttarakhand
(Dehra
Dun
, Lansdowne).
Andrewes (1933)
reported that the
holotype
was collected under bark of
Tectona grandis
, and several
paratypes
were in bamboo leaf roll, predaceous on
Hyblaea puera
(Cramer)
and
Hapalia machaeralis
Walker.