Neochya nitidissima (Chujo, 1935) comb. nov. Figures 9, 10

Luperodes nitidissimus Chûjô, 1935: 161; Chûjô 1962: 228.

Atrachya nitidissimus: Kimoto, 1966: 30 (additional records); Kimoto 1969: 55(additional records).

Luperodes bicoloripennis Chûjô, 1938: 138; Chûjô 1962: 227 (redescription). syn. nov.

Atrachya bicoloripennis: Kimoto, 1969: 55: (additional records); Kimoto 1986: 58 (additional records); Kimoto 1987: 190 (additional records).

Atrachya bicolor [sic!]: Kimoto, 1989: 257 (additional records).

Luperodes saramao Chûjô, 1962: 230. syn. nov.

Atrachya saramao: Kimoto, 1989: 257 (additional records).

Types.

Luperodes nitidissimus . Holotype ♀ (SDEI): "Kankau (Koshun = 恆春) / Formosa / H. Sauter V. 1912 [p, w] // Luperodes / nitidissimus / Chûjô [h] / DET. M. CHUJO [b, w] // Holotype [p, w, red letters]". Paratypes. 1♀ (TARI): "Formosa / Koshun, 1918 / IV 25-V 28 / J. Sonan, [p, w] // Allotype [h, w, red letters] // Luperodes / nitidissimus / Chûjô [h] / DET. M. CHUJO [b, w] // 2966 [p, w]"; 1♀ (TARI): "Formosa / Koshun, 1918 / IV 25-V 28 / J. Sonan, [p, w] // Paratype [h, w, red letters] // Luperodes / nitidissimus / Chûjô [h] / DET. M. CHUJO [b, w] // 2967 [p, w]".

Luperodes bicoloripennis . Lectotype ♂ (TARI, here designated): "Arisan (阿里山) / FORMOSA/ 24-25.V.1933 / Col. M. CHUJO [p, w] // CO / Type [p, w, circular label with yellow letters] // Luperodes / bicoloripennis / Chûjô [h] / DET. M. CHUJO [b, w] // 1383 [p, w]". Paralectotypes: 1♀ (TARI), same data as holotype but with “1382”; 1♂ (TARI): "Arisan [h] / FORMOSA [p] / 22.X.1931 [h] / COL. M. CHUJO [p, w] // CO / Type [p, w, circular label with yellow letters] // Luperodes / bicoloripennis / Chûjô [h] / DET. M. CHUJO [b, w] // 1381 [p, w]"; 1♀ (TARI), same but with “1373”; 1♂ (SDEI): "Arisan / FORMOSA / 24-25.V.1933 / COL. M. CHUJO [p, w] // Luperodes / bicoloripennis / Chûjô [h] / DET. M. CHUJO [b, w] // Syntypus [p, r]" 1♀ (SDEI), same but with "25.V.1933 [p]"; 1 (TARI, sex undetermined, abdomen lost): "ARISAN / XII.1915 [h] / Coll. M. Maki [p, w] // CO / Type [p, w, circular label with yellow letters] // Luperodes / bicoloripennis / Chûjô [h] / DET. M. CHUJO [b, w] // 1493 [p, w]"; 1♂ (TARI): "Formosa / Musha (霧社), 1919 / V 18-VI 15. / T. Okuni, [p, w] // CO / Type [p, w, circular label with yellow letters] // Luperodes / bicoloripennis / Chûjô [h] / DET. M. CHUJO [b, w] // 2587 [p, w]"; 1♀ (TARI), same but with “1492” . One specimen (♀, TARI) bearing type label: "Jujiro (十字路, in Chiayi) / 26-IV 1931 / Col. T. Shiraki [p, w] // CO / Type [p, w, circular label with yellow letters] // Luperodes / bicoloripennis / Chûjô [h] / DET. M. CHUJO [b, w] // 1494 [p, w]". It cannot be included in the type series since the data did not appear in the original description ( Chûjô 1938).

Luperodes saramao . Holotype ♂ (KUEC): "13 VIII 1936 / Mururoahu (給里洛山 =見晴山) / --- Kussya (庫霞 = 大同) / TAIHEIZAN (太平山) [p, pink label] // Luperodes / saramao / Chûjô [h, w] // Holotype [h, r]".

Other material.

Form A (N = 84). Chiayi: 1♂, 1♀ (TARI), Alishan (阿里山), 12.V.2011, leg. C.-F. Lee; 1♀ (TARI), same locality, 29.V.2016, leg. B.-X. Guo; 5♂♂, 1♀ (TARI), Shihshan channel (石山引水道), 23.XI.2013, leg. W.-C. Liao; 1♂, 2♀♀ (TARI), same locality, 21.I.2017, leg. W.-C. Liao; 8♂♂, 8♀♀ (TARI), Tzuchung (自忠), 21.IX.2009, leg. C.-F. Lee; Kaohsiung: 1♂ (TARI), Tienchih (天池), 17.V.2015, leg. B.-X. Guo; Nantou: 2♂♂ (TARI), Fenghuangshan (鳳凰山), 10.V.2010, leg. Y.-T. Wang; 2♂♂ (TARI), Hsitou (溪頭), 6.V.2009, leg. C.-F. Lee; 1♂ (NMNS), Shanlinchi (杉林溪), 10.V.1990, leg. C.-C. Chiang; 2♀♀ (NMNS), same locality, 19.V.1991, leg. C.-C. Chiang; 2♂♂, 3♀♀ (TARI), Tatachia (塔塔加), 9.VI.2009, leg. C.-F. Lee; 1♂, 1♀ (TARI), same locality, 21.VII.2009, leg. S.-F. Yu; 1♂, 1♀ (TARI), same locality, 21.IX.2009, leg. C.-F. Lee; 2♂♂ (TARI), same locality, 29.X.2009, leg. H. Lee; 1♂, 1♀ (TARI), same locality, 30.X.2009, leg. C.-F. Lee; 1♂, 2♀♀ (TARI), same locality, 17.XI.2009, leg. C.-F. Lee; 2♀♀ (TARI), same locality, 29.XII.2009, leg. M.-H. Tsou; 4♂♂, 4♀♀ (TARI), same locality, 17.XI.2009, leg. C.-F. Lee; 3♂♂ (TARI), same locality, 27.IV.2010, leg. C.-F. Lee; 1♂, 1♀ (TARI), same locality, 17.V.2010, leg. M.-H. Tsou; 2♂♂ (TARI), same locality, 13.VII.2014, leg. W.-C. Liao; Taichung: 1♂ (TARI), Anmashan (鞍馬山), 6-9.VII.1979, leg. L.-Y. Chou; 1♂ (TARI), same locality, 18.X.2008, leg. H. Lee; 2♂♂ (TARI), same locality, 15.X.2009, leg. J.-C. Chen; 1♀ (TARI), same locality, 7.VI.2010, leg. C.-F. Lee.

Form B (N = 255). Hsinchu: 1♀ (TARI), Kuanwu (觀霧), 30.IV.2009, leg. Y.-F. Hsu; 1♀ (TARI), same locality, 19.VIII.2009, leg. Y.-F. Hsu; 1♂ (TARI), same locality, 5.XI.2009, leg. H. Lee; 27♂♂, 7♀♀ (TARI), same locality, 30.IV.-1.V.2010, leg. C.-F. Lee & M.-H. Tsou; 1♂ (TARI), 3.VI.2011, leg. S.-F. Yu; 1♂ (TARI), Talulintao (大鹿林道), 17.II.2008, leg. M.-H. Tsou; 1♀ (TARI), same locality, 25.III.2009, leg. Y.-L. Lin; 2♂♂, 1♀ (TARI), same locality, 23.VI.2016, leg. Y.-L. Lin; Hualien: 2♂♂, 2♀♀ (TARI), Tayuling (大禹嶺), 2.VI.2016, leg. Y.-T. Chung; Miaoli: 1♂, 1♀ (TARI), Hsuehchien (雪見), 7.VI.2013, leg. W.-B. Yeh; 4♂♂ (TARI), Leshan (樂山), 14.VIII.2010, leg. H.-J. Chen; Nantou: 5♂♂, 3♀♀ (TARI), Huakung (華岡), 12.-13.IX.2010, leg. C.-F. Lee; 1♂ (TARI), same locality, 17.VI.2016, leg. J.-C. Chen; 1♂ (TARI), Lienhuachi (連華池), 23.-26.V.1980, leg. K.-S. Lin & B.-H. Chen; 3♂♂, 2♀♀ (TARI), Meifeng (梅峰), 10.V.1979, leg. K.-C. Chou; 2♂♂, 4♀♀ (TARI), same locality, 20.-22.VI.1979, leg. K.-S. Lin & B.-H. Chen; 1♂ (TARI), same locality, 23.V.-3.VI.1979; 1♀ (TARI), same locality, 18.VII.1979, leg. K.-C. Chou; 1♀ (TARI), same locality, 2.-12.X.1979; 1♂ (TARI), same locality, 24.X.1979, leg. K.-C. Chou; 1♂, 1♀ (TARI), same locality, 2.-4.VI.1980, leg. L.-Y. Chou & C.-C. Chen; 1♂, 1♀ (TARI), same locality, 5.-8.VI.1980, leg. C.-C. Chen; 2♂♂, 4♀♀ (TARI), same locality, 8.VI.1980, leg. K.-S. Lin & B.-H. Chen; 1♀ (TARI), same locality, 9.-16.VI.1980, leg. K.-S. Lin & B.-H. Chen; 6♂♂, 3♀♀ (TARI), same locality, 22.V.1982, leg. L.-Y. Chou; 3♂♂, 2♀♀ (TARI), same locality, 4.-7.X.1982, leg. K.-C. Chou; 1♂, 1♀ (TARI), same locality, 19-21.IV.1983, leg. K.-C. Chou & S.-P. Huang; 1♀ (TARI), same locality, 30.VII.1983, leg. L.-Y. Chou; 1♀ (TARI), same locality, 8.-11.V.1984, leg. K.-C. Chou & C.-C. Pan; 1♂ (TARI), same locality, 23.VII.1984, leg. K.-S. Lin; 1♀ (NMNS), same locality, 3.-15.X.1990, leg. C. K. Starr; 2♂♂ (NMNS), same locality, 27.II.1992, leg. Y.-C. Shiau; 1♀ (NMNS), same locality, 6.VIII.-11.IX.2001, leg. C.-S. Lin & W.-T. Yang; 5♀♀ (NMNS), same locality, 15.IV.-7.V.2003, leg. C.-S. Lin & W.-T. Yang; 2♀♀ (NMNS), same locality, 7.V.-11.VI.2003, leg. C.-S. Lin & W.-T. Yang; 1♂ (NMNS), same locality, 4.XI.-15.XII.2003, leg. C.-S. Lin & W.-T. Wang; 1♀ (NMNS), same locality, 6.IV.-11.V.2004, leg. C.-S. Lin & W.-T. Yang; 1♀ (NMNS), same locality, 5.X.-16.XI.2004, leg. C.-S. Lin & W.-T. Yang; 1♀ (NMNS), same locality, 12.IV.3.V.2005, leg. C.-S. Lin & W.-T. Yang; 1♀ (NMNS), same locality, 20.II.2009, leg. M.-L. Chan; 1♀ (TARI), same locality, 18.V.2009, leg. M.-H. Tsou; 2♀♀ (TARI), same locality, 15.IX.2009, leg. H. Lee; 2♀♀ (TARI), same locality, 15.IX.2009, leg. S.-F. Yu; 1♀ (TARI), same locality, 17.VI.2010, leg. C.-F. Lee; 2♂♂, 3♀♀ (TARI), same locality, 20.IV.2011, leg. C.-F. Lee; 1♂ (TARI), same locality, 30.V.2011, leg. M.-H. Tsou; 25♂♂, 18♀♀ (TARI), Peitungyanshan (北東眼山), 16.IX.2013, leg. F.-S. Huang; 2♀♀ (TARI), same locality, 28.III.2014, leg. F.-S. Huang; 3♂♂ (TARI), same locality, 14.IV.2014, leg. C.-F. Lee; 1♀ (NMNS), Piluchi (碧綠溪), 4.XII.1991, leg. Y.-C. Shiau; 1♂ (NMNS), same locality, 15.-31.III.1998, leg. M.-M. Yang; 3♂♂, 3♀♀ (TARI), Sungkang (松崗), 15.-17.VIII.1984, leg. K.-C. Chou; 1♂ (TARI), same locality, 18.IV.2015, leg. B.-X. Guo; 1♂, 1♀ (TARI), same locality, 10.IV.2016, leg. Y.-T. Chung; 1♂ (TARI), same locality, 2.VI.2016, leg. B.-X. Guo; 1♂, 3♀♀ (TARI), Tsuifeng (翠峰), IV.1984, leg. K.-S. Lin & K.-C. Chou; 1♂ (NMNS), Tunyuan (屯原), 29.IV.1992, leg. W.-T. Yang; 4♂♂, 1♀ (TARI), same locality, 18.X.2011, leg. J.-C. Chen; 1♂ (TARI), same locality, 30.IV.2017, leg. Y.-F. Hsu; 1♂ (NMNS), Yuanfeng (鳶峰), 5.VII.-2.VIII.2005, leg. C.-S. Lin & W.-T. Yang; Taichung: 9♂♂, 9♀♀ (TARI), Anmashan (鞍馬山), 6.-9.VII.1979, leg. L.-Y. Chou; 1♀ (NMNS), same locality, 1.V.1990, leg. C.-C. Chiang; 2♂♂ (NMNS), same locality, 3.V.1992, leg. C.-Y. Li; 1♂, 1♀ (NMNS), 22.IV.1998, leg. M.-M. Yang & H.-T. Chan; 1♀ (TARI), same locality, 16.VII.2007, leg. M.-H. Tsou; 1♀ (TARI), same locality, 7.VI.2010, leg. C.-F. Lee; 1♂ (TARI), same locality, 23.VII.2011, leg. J.-C. Chen; 9♂♂, 10♀♀ (TARI), Fushoushan (福壽山), 3.V.2016, leg. J.-C. Chen; 1♀ (NMNS), Nanhushi (南湖溪), 10.-11.VI.1988, leg. K.-W. Huang; 1♂ (TARI), Pilu (畢祿), 22.IV.2015, leg. C.-F. Lee; 1♀ (TARI), Wuleng (武陵), 27.-29.VI.1979, leg. K.-S. Lin & L.-Y. Chou; 1♂ (TARI), same locality, 6.IV.2014, leg. J.-C. Chen; 1♂ (TARI), same locality, 15.VIII.2014, leg. M.-H. Tsou.

Form C (N = 85). Hsinchu: 1♂ (TARI), Sumakusu (司馬庫斯), 26.IX.2009, leg. H.-J. Chen; Hualien: 1♀ (TARI), Kuanyuan (關原), 7.V.2006, leg. Y.-F. Hsu; 1♀ (TARI), same locality, 2.VII.2008, leg. M.-H. Tsou; 1♂, 2♀♀ (TARI), same locality, 2.VI.2016, leg. Y.-T. Chung & B.-X. Guo; 1♂ (NMNS), Kuanyun (觀雲), 13.V.2005, leg. J.-H. Chen; 1♂ (TARI), same locality, 19.VI.2010, leg. W.-P. Chan; 1♀ (TARI), Pilu (碧綠), 9.VII.2009, leg. C.-F. Lee; 1♀ (TARI), same locality, 10.IV.2014, leg. C.-F. Lee; 1♂ (TARI), same locality, 13.VI.2014, leg. C.-F. Lee; 1♀ (TARI), same locality, 23.IV.2015, leg. C.-F. Lee; 1♂ (TARI), same locality, 14.V.2015, leg. J.-C. Chen; 1♂, 1♀ (TARI), same locality, 7.VII.2015, leg. C.-F. Lee; 1♀ (TARI), same locality, 22.VII.2015, leg. U. Ong; 1♂, 2♀♀ (TARI), Sungyuan (松苑), 19.V.2018, leg. H.-F. Lu; 1♂ (TARI), Tayuling (大禹嶺), 6.-9.IX.1983, leg. L.-Y. Chou & K.-C. Chou; Kaohsiung: 2♀♀ (TARI), Chungchihkuan (中之關), 16.IV.2012, leg. L.-P. Hsu; 1♀ (TARI), same locality, 12.X.-6.XII.2012, leg. L.-P. Hsu; 1♂, 1♀ (TARI), same locality, 31.VII.2015, leg. C.-F. Lee; 1♂ (TARI), Tengchih (藤枝), 2.-5.VI.2008, leg. C.-F. Lee; 1♀ (TARI), same locality, 7.-10.XI.2008, leg. C.-T. Yao; 1♀ (TARI), same locality, 6.II.2009, leg. S.-F. Yu; 1♀ (TARI), same locality, 23.III.2009, leg. H. Lee; 1♀ (TARI), 26.V.2009, leg. C.-F. Lee; 1♀ (TARI), same locality, 6.VIII.2013, leg. B.-X. Guo; 1♀ (TARI), same locality, 8.III.2014, leg. W.-C. Liao; 3♂♂, 3♀♀ (NMNS), Tona (多納), 28.IV.1998, leg. M.-L. Chan; 1♀ (TARI), same locality, 2.VIII.2017, leg. B.-X. Guo; Miaoli: 1♂ (TARI), Hsuehchien (雪見), 12.III.2013, leg. W.-B. Yeh; 2♂♂ (TARI), same locality, 7.VI.2013, leg. W.-B. Yeh; 2♀♀ (NMNS), Taian (泰安), 19.XII.1989, leg. K.-W. Huang; Nantou: 1♂, Tunyuan (屯原), 12.VII.2014, leg. J.-C. Chen; Pingtung: 1♂ (TARI), Peitawushan (北大武山), 22.IX.2012, leg. J.-C. Chen; 1♂ (TARI), same locality, 25.VI.2018, leg. Y.-T. Chung; 1♀ (TARI), Tahanshan (大漢山), 20.VII.2007, leg. C.-F. Lee; 3♂♂, 8♀♀ (TARI), same locality, 6.II.2008, leg. M.-H. Tsou & S.-F. Yu; 3♂♂ (TARI), same locality, 3.III.2008, leg. C.-F. Lee; 1♀ (TARI), same locality, 4.VII.2008, leg. M.-H. Tsou; 3♂♂, 1♀ (TARI), same locality, 22.I.2009, leg. S.-F. Yu; 1♀ (TARI), same locality, 24.I.2009, leg. M.-H. Tsou; 1♂, 2♀♀ (TARI), same locality, 5.IV.2009, leg. C.-F. Lee; 1♀ (TARI), same locality, 26.XI.2009, leg. J.-C. Chen; 1♀ (TARI), 15.II.2010, leg. M.-H. Tsou; 1♂ (TARI), same locality, 14.IV.2011, leg. J.-C. Chen; 1♀ (TARI), same locality, 14.XII.2011, leg. J.-C. Chen; 1♀ (TARI), same locality, 19.XI.2012, leg. J.-C. Chen; 1♀ (TARI), same locality, 26.III.2013, leg. C.-F. Lee; 1♀ (TARI), same locality, 3.IV.2013, leg. Y.-T. Chung; 3♂♂ (TARI), same locality, 17.III.2014, leg. Y.-T. Chung & J.-C. Chen; 1♂ (TARI), same locality, 2.III.2015, leg. Y.-T. Chung; 1♀ (TARI), same locality, 24.III.2017, leg. Y.-T. Chung; Taichung: 1♂ (TARI), Pilu (畢祿), 18.VI.2010, leg. C.-F. Lee; 1♂ (TARI), same locality, 2.VI.2016, leg. J.-C. Chen.

Form D (N = 44). Chiayi: 4♂♂, 1♀ (TARI), Laichitashan (來吉塔山), 19.III.2009, leg. H. Lee; Hsinchu: 1♂, 1♀ (TARI), Leetungshan (李棟山), 15.III.2009, leg. S.-F. Yu; 4♂♂, 1♀ (TARI), same locality, 27.X.2009, leg. S.-F. Yu; 1♀ (TARI), Mamei (馬美), 4.V.2008, leg. S.-F. Yu; 1♀ (TARI), Taikang trial (泰崗林道), 8.IX.2013, leg. Y.-L. Lin; Ilan: 1♀ (TARI), Suyuan (思源), 15.IV.2009, leg. M.-H. Tsou; 1♀ (TARI), same locality, 9.VI.2009, leg. S.-F. Yu; 2♀♀ (TARI), same locality, 12.IX.2010, leg. M.-H. Tsou; 1♂, 1♀ (TARI), same locality, 11.VIII.2014, leg. H. Lee & J.-C. Chen; 1♂ (TARI), same locality, 30.VII.2015, leg. H. Lee; 1♀ (TARI), Taipingshan (太平山), 13.VI.2007, leg. Y.-C. Chang; Pingtung: 1♀ (TARI), Kenting (墾丁), 23.VIII.2016, leg. Y.-T. Chung; Tainan: 1♂ (TARI), Kantoushan (崁頭山), 17.VI.2012, leg. W.-C. Liao; 1♂, 1♀ (TARI), Meiling (梅嶺), 28.XII.2008, leg. U. Ong; Taipei: 1♂, 1♀ (TARI), Fushan (福山), 19.VI.2007, leg. M.-H. Tsou; 1♂ (TARI), Houtung (侯硐), 25.V.2009, leg. J.-C. Chen; Taoyuan: 1♂ (TARI), Lalashan (拉拉山), 7.VIII.2008, leg. H.-J. Chen; 1♂, 1♀ (TARI), same locality, 30.X.2008, leg. S.-F. Yu; 2♂♂, 2♀♀ (TARI), same locality, 8.III.2009, leg. S.-F. Yu; 1♂, 1♀ (TARI), same locality, 2.IV.2009, leg. C.-F. Lee & H.-J. Chen; 4♂♂, 1♀ (TARI), same locality, 14.V.2009, leg. C.-F. Lee; Yunlin: 1♀ (TARI), Shihpishan (石壁山), 26.IV.2015, leg. W.-C. Liao.

Form E (N = 1). Pingtung: 1♂ (TARI), Peitawushan (北大武山), 25.VI.2018, leg. Y.-T. Chung.

Redescription.

Length 5.6-7.0 mm, width 2.9-3.8 mm. General color extremely variable, with five distinct color patterns (see variation). Antennae (Fig. 10A) filiform in males, ratio of length of antennomeres I to XI 1.0: 0.4: 0.4: 1.0: 1.1: 1.1: 1.1: 1.0: 1.1: 0.9: 1.0; ratio of length to width from antennomere I to XI 3.6: 1.6: 1.8: 4.1: 5.0: 5.3: 5.9: 5.1: 5.9: 5.2: 5.4; similar in females, ratio of length of antennomeres I to XI (Fig. 10B) 1.0: 0.4: 0.4: 1.0: 1.1: 1.0: 1.1: 1.0: 1.0: 0.9: 1.0; ratio of length to width from antennomere I to XI 3.6: 1.8: 2.1: 4.5: 5.3: 5.2: 5.4: 5.5: 5.5: 5.0: 5.8. Pronotum 1.90-1.94 times wider than long; lateral margins slightly rounded, basal margin slightly rounded, apical margin slightly concave; disc with dense minute punctures, but without lateral depressions. Elytra 1.36-1.40 times longer than wide; lateral margins rounded, widest at middle; disc moderately convex, with dense, minute punctures; apex truncate. Penis (Fig. 10C-E) wide, ca. 3.8 times longer than wide; lateral margins rounded, widest at basal 1/3; apex broadly rounded; tectum elongate from apical 1/6 to middle, parallel-sided, apex broadly rounded; slightly and apically curved in lateral view; ventral surface with membranous area from apex to apical 1/4; with one small rounded process on lateral margin at apical 1/4. Endophallic spiculae complex with six or seven pairs of hooked spiculae (visible in dorsal view), one additional pair of hooked spiculae near middle with four or five ventral branches; with one pair of longitudinal rows of hair-like setae and small rounded sclerites near base. Gonocoxae (Fig. 10F) slender, tightly conjunct from apical 1/6 to middle; each gonocoxa with eight setae from apical 1/6 to apex, subapically widened, apex narrowly rounded, base deeply bifurcate. Ventrite VIII (Fig. 10G) weakly sclerotized except apex, with several short and long setae at apex, and several long setae at sides, spiculum elongate. Spermathecal receptaculum (Fig. 10H) as slender as pump, apically tapering; pump slender and curved; sclerotized spermathecal duct extremely elongate, but base wide, followed by slender tube, then with inflated areas. Bursal sclerites reduced.

Variation.

Color pattern divided into four forms. Form A (Fig. 9A, B) (described as one form of Atrachya bicoloripennis): general color black; but apical 2/3 white, abdomen reddish brown. Form B (Fig. 9C, D): similar to form A, but white area replaced with red (described as another form of A. bicoloripennis). Form C (Fig. 9E, F) (described as A. saramao): similar to form B, but elytra entirely reddish brown, meso-and metathoracic ventrites reddish brown; some individuals have paler femora and antennae. Form D (Fig. 9G, H) (described as typical form of A. nitidissima): body color reddish brown, but antennae, tibiae, and tarsi darker. Form E (Fig. 9I): similar to form A, but apical 1/3 of elytra black.

Diagnosis.

Neochya nitidissima ( Chûjô) is similar to N. chengi sp. nov. in having wide elytra, truncate elytral apices and reduced lateral depressions on the pronotum (Figs 4, 9) (narrow elytra, rounded elytra apices and with lateral depression on the pronotum in others (Figs 2D-F, 7) but differs from N. chengi sp. nov. in rounded elytra and having reduced punctures on the pronotum and fine punctures on the elytra (Fig. 9) (parallel sided elytra and coarse punctures on pronotum and elytra in N. chengi sp. nov. (Fig. 3)). In addition, males of both species are separated from others with smooth margin of tectum of the penis (Figs 5C, 10C) (serrate margin of tectum (Figs 8C, 12C), but males of N. nitidissima differs from those of N. chengi with small rounded process on lateral margin of the penis (Fig. 10C-E) (lacking small rounded process on lateral margin of the penis in N. nitidissima (Fig. 5C-E). Atrachya bicoloripennis (Forms A and B: Fig. 9A-E) and A. saramao (Form C: Fig. 9E, F) have distinct color patterns which are different from typical form N. nitidissima (Form D: (Fig. 9G, H)). All of they are synonyms with no doubt based on examination of the penis.

Remarks.

The holotype of Luperodes nitidissimus was described as a male ( Chûjô 1935), but it is actually a female.

Food plants.

Celastraceae: Celastrus kusanoi Hayata, C. hindsii Benth., Euonymus spraguei Hayata (Fig. 1D).

Distribution.

Widespread in Taiwan. Most individuals with different color patterns can be separated based on distributions (Fig. 11A) except for the single form E. Members of form A occur at high elevation (> 2000 m) in south Taiwan, including Chiayi, south Nantou, and Kaohsiung counties. Those of form B also occur at high elevation (> 2000 m) but in central Taiwan, including Hsinchu, Miaoli, north and central Nantou, Taichung and Hualien counties. Those of form C occur at middle elevations (1000-2000 m) in central and south and east Taiwan, including Hsinchu, Miaoli, Taichung, Nantou, Kaohsiung, Pingtung, and Hualien counties. Those of form D occur at middle elevations (1000-2000 m) in north Taiwan, including Taipei, Taoyuan, Hsinchu, and Ilan counties, as well as lowlands (<1000 m) in central and south Taiwan, including Yulin, Tainan, and Pingtung counties. Coexistence of two different color patterns was observed in some areas, such forms A and B at Anmashan (鞍馬山); forms B and C at Hsuehchien (雪見), Tayuling (大禹嶺), and Tunyuan (屯原); forms C and E at Peitawushan (北大武山).