Onycholyda nigroclypeata Shinohara, 1987

(Figs 37–39) (https://doi.org/10.6084/m9.figshare.11405073)

Pamphilius tenuis: Kim, 1970: 126, 717; Kim, 1980: 3. Not Takeuchi, 1938.

Onycholyda nigroclypeata Shinohara, 1987b: 649; Shinohara, 1992a: 502; Shinohara & Byun, 1993: 84; Kim et al., 1994: 216; Zhelochovtsev & Zinovjev, 1995: 398; Shinohara & Lee, 1997: 215; Shinohara, 2002b: 421; Shinohara, 2004: 262; Shinohara & Lelej, 2007: 929; Shinohara & Taeger, 2007: 32; Paek et al., 2010: 161; Taeger et al., 2010: 84; Shinohara & Lee 2011: 209; Sundukov & Lelej, 2012: 108; Sundukov, 2017: 103; Lee et al., 2019: 8; Shinohara & Tripotin, 2021a: 60; Shinohara & Tripotin, 2021b: 196.

Material examined. Forty specimens, including the type series. Thirty-nine specimens are from the Russian Far East and North and South Korea (Shinohara 1987b; Shinohara & Byun 1993; Shinohara & Lee 1997; Shinohara & Taeger 2007; Shinohara & Tripotin 2021a, b; present work). New collection data: RUSSIA: Primorskij Kraj: 1♀ (Fig. 38, DEI-GISHym 86743), Partizansk SSW, 140m, 43.073°N 133.074°E, 9. VI. 2016, K. Kramp, M. Prous & A. Taeger, RU044 (SDEI). SOUTH KOREA: Gangwon-do: 2♀ 1♂, Mirugam (Bukdaesa), 1,300 m, Odaesan Mts., 29. V.–1. VI. 1998, A. Shinohara (NSMT) ; 1♀, same locality, 2. VI . 2002, A. Shinohara (NSMT).

Distribution. Russia (Khabarovskij Kraj, Primorskij Kraj), North and South Korea, China (Heilongjiang) (Shinohara 1992a).

Host plant. Rosaceae: Agrimonia pilosa Ledeb. (Shinohara & Lee 2011).

Remarks. This species has much in common with O. decorata Shinohara, 1985, from Japan, in adult and larval morphology and host plant (Shinohara 1987b; Shinohara & Lee 2011). In our molecular analyses, the two species were also quite close, diverging by a minimum of 1.1% in COI and 0.3% in NaK. The two species formed a clade with 100% UFBoot support in both COI and NaK analyses (Figs 139, 153). However, we treat them as distinct species, because the differences between them (Shinohara 1987b) are small but quite constant.

The newly acquired female specimen from Russia (Fig. 38) has extensively orange-marked temples and vertex, but is otherwise not distinguishable from other specimens (Fig. 37) (Shinohara 1987b). The pale coloration of the head of this specimen is asymmetrical suggesting its aberrant nature (Fig. 38e).