Hecullus bracteatus (Ball)

(Figs. 44–47, 94Q–R)

Hecalus bracteatus Ball, 1901:4 [original description]; Crumb, 1911: 234 [distribution]; Van Duzee, 1916: 69 [distribution]; Lawson, 1920: 117 [description]; Beamer, 1937: 12 [morphology]; DeLong & Knull, 1946: 26 [distribution]

Hecullus bracteatus: Oman, 1949 a: 31 [listed, classification]; Hamilton, 2000: 434–436 [morphology, illustrated as H. balli]; Zahniser & Webb, 2004: 673 [illustrated]; Zahniser & Dietrich, 2015: 479, 482 [phylogeny, DNA sequences]; Zahniser, 2007 [online catalogue]

Diagnosis. H. bracteatus can be distinguished from other Hecullus spp. by its size (male 4.3–5.0 mm, female 6.0– 7.0 mm), female head length 1.3x or less interocular width, male forewings with brown shading along vein margins, male head and pronotum with distinct striping, aedeagus in lateral view with apex attenuated to sharp apex, and ventral processes not projecting caudad and closely following curvature of aedeagus.

Material examined. 3♂, 7♀: CO: Rocky Ford, 16-VI-1900 [USNM] . 1♀: AZ: Tombstone, 1-VIII-1931 [USNM] . 7♂, 5♀, 2n: AZ: Ash Fork, 16-VIII-1929 [USNM] . 8♀: AZ: Chiricahua Mts., 4-VII-1930 [UNSM] . 1♂: NM: Espanola, 18-VI-1938 [USNM] . 1♂: UT: Kanab, 6-VIII-1939 [USNM] . 3♀: NM: Hurley, 31-VII-1935, R.H. Beamer [USNM] . 2♀, 1n: AZ: Huachuca Mts., 31-VII-1935, E.D. Ball [USNM]. 1n: AZ: Bisbee, 4-IX-1936, E.D. Ball [USNM] . 2♀: AZ: Benson, 9-VIII-1937, E.D. Ball [USNM] . 1♀: AZ: Painted Desert, 28-VIII-1934 [USNM] . 1♂: AZ: Tucson, 24-IV-1936 [USNM] . 1♀: NV: Las Vegas, 8-IX-1933, E.W. Davis [USNM] . 1♀: AZ: Wilcox [USNM] . 1♀: CO: Las Animas, 17-VII-1901 [USNM] . 1♂, 4♀, 12n: NV: Overton. 1♀: UT: Orderville, 7-VIII- 1930 [USNM] . 1♀: AZ: Far Away R., 12-IX-1935 [USNM] . 1♂: NM: Moses, 21-VI-1984 [USNM] . 1♂: NM: Hidalgo Co., Indian Ck. nr. Culberson Camp, 31°22’28”N 108°36’25”W, 7–8-VIII-2002, J. George, PT, UCRC ENT 154896 [UCRC] . 4♀: AZ: Painted Desert, “ert 1.IX.1932 ” . 1♂, 1♀: AZ: Naco, 27-IV-1932 [USNM] . 1♀: AZ: Nogales, “Sept.”, A. Koebele [USNM] . 42♀: AZ: Pearce, 4-VII-1930, E.D. Ball [USNM] . 1♂, 1♀: NM: Laguna, 22-V-2002, 35.03797°N 107.35566°W, J.N. Zahniser #02-008 [USNM] . 1♂, 1♀, 1n: NM: Harding Co., Kiowa Na- tional Grassland, Mills Canyon Creek Campground, sweep, 3–4-IX-2009, JN Zahniser #09-21 [USNM] .

Distribution. H. bracteatus is known from the United States throughout Arizona and New Mexico and in the southern parts of Nevada, Utah, and Colorado.

Remarks. This is the most well-known species of the genus and has a significantly wider known geographic range than H. balli (Fig. 47). However, more collecting effort in northern Mexico may show wider distributions for these species and may reveal undescribed species. The male genitalia are illustrated here for the first time. Illustrations stated to be of H. balli by Hamilton (2000; Figs. 7, 14) appear to be of H. bracteatus . Specimens from New Mexico (Laguna and Kiowa National Grassland) were swept or vacuumed from grasses.