Physalaemus signifer clade

Species in this clade usually have A calls with low fundamental frequency, below 500 Hz (except P. bokermanni; see below). Usually the fundamental band is absent or with very low energy, barely visible in audiospectrograms (all species except some calls of P. irroratus, P. obtectus, and P. nanus). Most of the species (66.6 %) have more than one call type. Nonlinear phenomena, subharmonics and deterministic chaos, are common in the clade (see P. angrensis, P. atlanticus, P. bokermanni, P. camacan, P. crombiei, P. irroratus, P. moreirae, P. nanus, P. obtectus, P. signifer, and P. spiniger) even though usually restricted to parts of some calls (i.e., polymorphic in presence and position; vs. present in all calls in the same call positions in species of P. cuvieri clade). Eight species ( P. angrensis, P. atlanticus, P. bokermanni, P. crombiei, P. irroratus, P. moreirae, P. nanus, and P. obtectus) have calls with strong PAM, with silence periods between pulses.