Europharinodes Yin & Cai gen. nov.

Figs 2, 3, 4, 5

Type species.

Europharinodes schaufussi sp. nov.

Diagnosis.

Most of body covered with squamous setae; body generally compact; head, pronotum and femora coarsely punctate. Head roundly-triangular, with short and narrow frontal rostrum; large vertexal and frontal foveae present; antennal insertions close, antennomeres slightly elongate to sub-moniliform, clubs formed by apical three enlarged antennomeres; lacking ocular-mandibular carinae; clypeus sharply sloping, anterior margin carinate and rounded, lateral margins straight to eyes; maxillary palpi small, four-segmented, with short apical palpal cone. Pronotum with median and lateral antebasal foveae obscured by squamous setae; with antero-hypomeral foveae, lacking hypomeral carinae. Elytra each with two large basal foveae and two distinct discal striae. Each tarsus plesiomorphically with two subequal claws. Abdomen with tergite 1 (IV) slightly longer than 2 (V), tergites 1-4 (IV-VII) broadly sulcate at bases; paratergites moderately broad and laterally protruding. Aedeagus symmetrical, basal capsule enlarged, paired parameres elongate.

Description.

Habitus (Figs 2B-D; 3A, B) stout, compact; most of body with squamous setae. Head (Fig. 3E) roundly triangular; vertex coarsely punctate, vertexal foveae (= dorsal tentorial pits; Fig. 3E; vf) relatively large, obscured by squamous setae; frontal fovea (Fig. 3E; ff) distinct, rostrum short and narrow, antennal insertions located at ventral surface of rostrum, tubercles barely raised; compound eyes (Fig. 3G; ce) large, prominent; lacking ocular-mandibular carina; clypeus (Fig. 3G; cl) sharply sloping, with broadly rounded, ridged anterior margin; labrum (Fig. 3G; la) transverse, subtrapezoidal; mandible (Fig. 3G; ma) with single distinct apical and preapical teeth; maxillary palpus small (Fig. 3H, I), four-segmented, with palpomere 1 small, indistinct, 2 (Fig. 3H; p2) pedunculate basally and broadened toward apex, 3 (Fig. 3H; p3) roundly triangular, 4 (Fig. 3H; p4) sub-fusiform, with short palpal cone at apex. Venter with small, widely separated gular foveae (= posterior tentorial pits; Fig. 3F; gf) in shared transverse impression, gular moderately raised along middle, weakly impressed admesally; neck region broad. Paired tentorial arms (Fig. 3E; ta) V-shaped, each branched at base and extending anteriorly to reach inner wall of clypeus (right arm broken at apex in holotype). Pharynx (Fig. 3E; ph) about 2/3 of head length, posteriorly broadened. Antenna 11-segmented, moderately elongate, extending to approximately half of elytral length when extending posteriorly, antennomeres each slightly elongate to sub-moniliform, club (Fig. 3F) formed by enlarged apical three antennomeres.

Pronotum (Fig. 4A) approximately as long as broad, sides rounded, broadest at middle, anteriorly and posteriorly narrowed, surface with dense squamous setae (Fig. 3C); disc coarsely punctate, with squamous setae all over surface, covering median and lateral antebasal foveae (Fig. 4A, B; maf, laf), lacking sulcus or carina. Hypomeron extended, lacking carina, with anterior hypomeral fovea (Fig. 4B, C; ahf). Prosternum (Fig. 4C) with widely separated lateral procoxal foveae (Fig. 4C; lpcf), anterior part shorter than coxal part, margins of coxal cavity moderately carinate.

Elytra broadly truncate at bases, each elytron with two large basal foveae (Fig. 4D; bef), with two long discal striae (Fig. 4G; ds) extending from foveae to posterior elytral margin, lacking subhumeral fovea or marginal sulcus. Metathoracic (hind) wings (Fig. 2D; hw) fully developed.

Mesoventrite with median foveae (Fig. 4E; mmsf) originating from shared transverse impression, moderately separated, with pairs of large lateral and distinct anterolateral mesoventral foveae (not shown in figure); metaventrite with large, setose lateral coxal foveae (Fig. 4F; lmcf), with single median metaventral fovea (Fig. 4E; mmtf), posterior margin with small, narrow split at middle (Fig. 4E; ms); metacoxae broadly separated. Marginal carina of meso- and metaventrite complete (Fig. 4F; mc).

Legs (Fig. 5A-C) moderately elongate; femora roughly punctate; with short tarsomere 1 and long tarsomere 2 and 3, 2 slightly longer than 3, each tarsus with two subequal pretarsal claws (Fig. 2G).

Abdomen constricted at base, with dense squamous setae (Fig. 3D); tergite 1 (IV) slightly longer than 2 (V), 2-4 (V-VII) subequal in length, tergites 1-4 (Fig. 4G, H; t1-4) each deeply and broadly sulcate at base, at lease with one pair of basolateral foveae (Fig. 4G; blf), tergite 5 (VIII) (Fig. 4H; t5) roundly triangular, transverse, medially roundly emarginate at posterior margin; sternite 2-5 (IV-VII) (Fig. 4H, I; s2-5) each broadly sulcate at base, with one pair of basolateral foveae (Fig. 3H; blf), 6 (VIII), sternite 6 (Fig. 4H, I; s6) transverse.

Males have modified sternites 4 and 5 (VI and VII). Aedeagus (Fig. 5D-F, I-K) dorso-ventrally symmetrical.

Etymology.

The new generic name is a combination of Latin ' Eurōpa (Europe)' and genus Apharinodes, referring to the origin of the fossil in Baltic amber and its affinity with Apharinodes . The gender is feminine.