Isatis velutina Boiss. & A. Huet in Boiss., Diagn. Pl. Orient. ser. 2, 5: 45. 1856.

Type: “Hab. in valle Kassuklu Armeniae borealis cl. Huet du Pavillon”.

Holotypus: TURKEY: “In valle Kassuklu”, V.1853, Huet du Pavillon s.n. (G-BOIS [G00332595]; iso-: BM [BM001254135], G [G00371803], K [K000642859]) .

Notes. – The isotype sheet at G has a label with the handwritten part “In ruderatis vallis Kassuklu 4–5000 p.s.m.” and a printed part “Inter Baibout et Erzeroum. Armenia ”. The sheet was erroneously annotated by Davis as holotype .

As discussed under I. candolleana, the fate of I. velutina depends on whether the mixed type collection of the lectotype of the former is treated as one or two species. It is concluded here that two species are involved, though DAVIS (1965b: 296), followed by GREUTER et al. (1986: 129), reduced I. velutina to synonymy of I. candolleana and accepted the latter as a polymorphic species with two fruit types.

Davis annotated the holotype sheet of I. velutina in 1960 (Fig. 32, p. 124) as a synonym of the later-published I. conringiifolia Bornm., but he (DAVIS, 1964: 21, 1965b: 296) treated both as synonyms of I. candolleana . He also annotated the isotype G00371803 as holotype of I. velutina and “probably best treated as a var. of I. candolleana, or possibly a hybrid between the latter & sp. aff. I. atropatana Grossh. ”.