Hesperis unguicularis Boiss., Diagn. Pl. Orient. ser. 2, 5: 21. 1856.

Type: “Hab. in Armeniâ cl. Calvert”.

Holotypus: TURKEY: “Erzeroum”, 1853, Calvert 93 (G-BOIS [G00332219]; iso-: JE [JE00002543]) .

= Hesperis bicuspidata (Willd.) Poir., Encycl. Suppl. 3: 195. 1813.

Notes. – It seems that only the above two duplicates are available for H. unguicularis in the major herbaria consult-ed. Although DVOŘÁK (1965: 416), correctly indicated that the holotype is at G (for G-BOIS) and isotype is at JE, he (DVOŘÁK, 1973a: 117) reversed action by indicating that the type, rather than isotype, is at JE. Boissier did not examine or annotate the JE sheet, and therefore it makes perfect sense to treat the G-BOIS material as the holotype.

DVOŘÁK (1965, 1973b) recognized the species as a distinct instead of reducing it to synonymy of H. biscuspidata, but the differences between them are trivial, and it is more sound to recognize the latter as a variable species accommodating the former.