Hydrobiosella Tillyard 1924

Hydrobiosella Tillyard 1924: 288; Mosely and Kimmins 1953: 387; Neboiss 1977: 45; Neboiss 2003: 55.

Type species. Hydrobiosella stenocerca Tillyard by monotypy.

Generic descriptions are given by Tillyard (1924: 288); Mosely and Kimmins (1953: 387) and Neboiss (1977: 45).

Key to males of known Australian groups (or ungrouped species) of Hydrobiosella Tillyard (after Cartwright 2010, 2012)

1. Phallus without pair of parameres (Figs 2, 3, 5, 6; Cartwright 2012a, figs 2, 3; Neboiss 1986, figs pp. 99, H. amblyopia; 101, H. tasmanica; 102, H. corinna) 2

- Phallus with pair of parameres (Cartwright 2010, figs 2, 3, 5, 6; Neboiss 1986, figs pp. 99, H. michaelseni, H. waddama; 101, H. letti; 102, H.bispina; 103, H. arcuata) 6

2. Preanal appendages present, usually small (Cartwright2012a, figs 2, 3; Neboiss 1977, figs 204, 205, 216, 217; Neboiss 1986, figs pp. 101, H. tasmanica; 102, H. corinna; Neboiss 2003, figs 8a–h) 3

- Preanal appendages absent (Figs 2, 3, 18, 19; Neboiss 1986, figs pp. 99, H. amblyopia; 101, H. tasmanica) 4

3. Preanal appendages relatively slender, elongate and ‘unattached’ to segment IX (Cartwright 2012a, figs 2, 3, 5); NE-Qld Hydrobiosella eminentia group

- Preanal appendages often short and bulbous or ‘attached’ to segment IX (Neboiss, 1977, figs 204-211; Neboiss 1986, figs p. 102, H. corinna; Neboiss, 2003, figs 8A-H); Tas Hydrobiosella corinna group

4. Phallus apically with downward projecting spine(s) (Neboiss 1977, figs 216-221, 225, 226; Neboiss 1986, figs p. 101, H. armata, H. tasmanica; Neboiss 2003, figs 10A-G, 11A-G, 12A-F); Tas H. tasmanica group

- Phallus apically without downward projecting spine(s) (Figs 2, 3; Neboiss 1982, fig. 12; Neboiss 1986, figs p. 99 H. amblyopia) 5

5. Segment IX with a large disto-lateral projection (Neboiss 1982, fig. 12; Neboiss 1986, figs p. 99 H. amblyopia); S-WA H. amblyopia (ungrouped)

- Segment IX without a large disto-lateral projection (Figs 2, 3, 5, 6); C and NE Vic H. bandabanda group

6. Inferior appendages with harpago with dark row of setae forming fringe along ventral margin (Cartwright 2010, figs 3, 6; Neboiss 1986, figs pp. 102, H.bispina; 103, H. arcuata); E-Vic, E-NSW, E-Qld Hydrobiosella bispina group

- Inferior appendages with harpago without dark row of setae forming fringe along ventral margin (Figs 18, 19; Neboiss 1986, figs pp. 99, H. michaelseni, H. waddama) 7

7. Parameres elongate and sinusoidal, attached ventrally to base of phallus (Cartwright 2012b, figs 2, 3, 5, 6; Neboiss 1977, fig. 233; Neboiss 1986, figs p. 99, H. waddama); Tas, SE Aust. Hydrobiosella waddama group

– Parameres not elongate and sinusoidal, not attached ventrally to base of phallus (Figs 18, 19; Neboiss 1982, figs 9, 10; Neboiss 1986, figs pp. 99, H. michaelseni; 101, H. letti) 8

8. Parameres curved strongly and crossed (Neboiss 1982, figs 9, 10; Neboiss 1986, figs p. 99; H. michaelseni); S-WA Hydrobiosella michaelseni (Ulmer) (unplaced to group)

- Parameres not curved strongly and crossed (Figs 18, 19; Neboiss 1986, figs p. 101, H. letti); NE-Vic, CE-NSW, E-Qld Hydrobiosella letti group