Jujubinus exasperatus (Pennant, 1777) (Fig. 4F 1-F 3)

Trochus exasperatus Pennant, 1777: 126 .

Trochus (Calliostoma) exasperatus – Harmer 1923: 723, pl. 58, fig. 18.

Jujubinus (Jujubinus) exasperatus – Cuerda Barceló 1987: 185, pl. 15, figs 11, 12.

Jujubinus exasperatus – Landau et al. 2003: 42, pl. 9, fig. 6. — Chirli 2004: 78, pl. 32, figs 10-12, pl. 33, figs 1-4. — Chirli & Linse 2011: 39, pl. 6, fig. 1. — Chirli & Forli 2017: fig. 6b.

Jujubinus aff. exasperatus – Landau et al. 2017 97, pl. 21, figs 1-4.

For more, see synonymy list in Landau et al. (2003).

MATERIAL AND DIMENSIONS. — Maximum height 9.4 mm, width 7.3 mm. RGM.1364903 (36), leg. WG; RGM.1364969 (48), leg. ACJ; RGM.1365178 (1), leg. ACJ; RGM.1365067 (60+) leg. AWJ .

SPECIES CHARACTERISATION. — Shell trochiform; protoconch severely worn or not preserved. Teleoconch of six flat-sided whorls separated by superficial suture; sculpture of five spiral cords, abapical one broader, stronger, placed just above the suture; fine lamellar ribs visible in cord interspaces giving them a pitted appearance; base flattened bearing six concentric cords; aperture subquadrate; columella short, slightly oblique bearing subobsolete fold.

DISTRIBUTION. —?Upper Miocene: NW France (Brébion 1964; conspecificity uncertain; Landau et al. 2017). — Pliocene: Italy (Brunetti & Vecchi 2005). — Upper Pliocene. Western Mediterranean, S. Spain (Landau et al. 2003). — Lower Pleistocene: Atlantic, southern England (Harmer 1923), Selsoif, France (this paper); central Mediterranean, Italy (Chirli & Forli 2017); eastern Mediterranean, Rhodes Island (Chirli & Linse 2011). — Upper Pleistocene: Atlantic, Santa Maria Island, Azores (Callapez & Ferreira 2000); western Mediterranean, Balearic Islands (Cuerda Barceló 1987). Today, this species occurs in the Atlantic from the west coast of Scotland to the Canary Islands and Azores (Fretter & Graham 1977; Hernández et al. 2011), Mediterranean Sea (Manousis 2021) and Aegean Sea (Öztürk et al. 2014).

REMARKS

Jujubinus exasperatus (Pennant, 1777) is a highly variable species that is best distinguish from other Jujubinus species by regular spiral cords with a stronger abapical cord.