Tentyria velox ssp. circumvoluta nom. nov. and stat. nov. (Figs. 29, 63, 99, 134, 215, 245)

Tentyria subrugosa Solier 1835: 352 nom. preocc., homonym nov.; Kraatz 1865: 132, Reitter 1900: 171, Fuente 1934: 30, Español 1960: 408, Viñolas & Cartagena 2005: 82, 357d, L̂bl & Smetana 2008: 206, Martínez 2018: 58, Iwan & L̂bl 2020: 252.

Non Tentyria subrugosa Besser 1832:13 (= Tentyria italica Solier 1835: 348 syn. nov.).

Types examined: Tentyria subrugosa ?? Sol., Espag. Du64 (Dupont 64, circular label) / MUSEUM PARIS Coll. Solier, COLL. DE MARSEUL 2842-90 / Tentyria subrugosa Sol., det. J. Ferrer (1♀, MNHN) (Fig. 245).

Five specimens from the NMHUB, one of them with the following labels: subrugosa Dj. S. Par; Sicil. (old handwritten label) / (modern label) Hist. Coll. ( Coleoptera), Nr. 45571, Tentyria subrugosa Dej ?, Sicil. Parreyss, Zool. Mus. Berlin. The others four specimens carrying similar modern labels, and one of them, with smooth elytra, also bears one old handwritten label, similar to the first: Tentyr. subrugosa, Dej ?, Sicil. Parr., and another old label with the reference number “45571”.

Aditional material: Ávila: La Serrada, ÁVILA, Spain, V-1960 / J. Vives leg. / Tentyria subrugosa Sol. (1 ♁, CJLB and 2 exx, CJF); Ávila, J. Sanz / T. subrugosa Sol. / MNCN_Ent, Nº Cat. 70795 / Tentyria subrugosa Sol., det. J.L. Bujalance (1♁, MNCN); Ávila, T. subrugosa Sol. / MNCN_Ent., Nº Cat. 70871, Tentyria subrugosa Sol., J.L. Bujalance det. (1♀, MNHN).

Diagnosis: Body ovate, elongate (Fig. 215), integument black, not very shiny. Head (Fig. 29) elongate in front of the eyes, which are large and moderately convex, the head is also provided with dense punctures, especially in the front part; the supraorbital folds much enhanced, delimiting two large depressions inwards, at the level of the insertion of the antennae; epistome subtruncate or rounded, projecting forward and provided with a very small, hardly perceptible tooth; gular groove (Fig. 63) transverse, narrow, except in the middle, and moderately deep. Prothorax transverse, slightly convex, with fine but dense points, slightly arched at its anterior edge, sides rounded and somewhat narrower posteriorly; base bisinuate before the rear angles and prolonged in curve posteriorly, with a broad marginal groove and thickened rim (Fig. 99); prosternal apophysis (Fig. 134) parallel-sided, depressed in the middle, with the end slightly bent upwards, giving it a rounded appearance in ventral view. Elytra elongate oval, moderately convex, with inconspicuous punctures, provide with sinuous and irregular folds, as longitudinal as transversal, and some suborbicular tubercles, base slightly arched and with moderately thick and full rim. Females show less elongated oval elytra and the base of the pronotum less prolonged backwards, generally with a coarser appearance.

T. velox ssp. circumvoluta differs from T. velox Chevrolat by its somewhat larger average size, the shape of the head, which is more elongated in front of the eyes, the base of the pronotum more sinuate and curved, and particularly by the surface of the elytra, provided with tuberous wrinkles remembering cerebral convolutions, and by the shape more ovate-elongated particularly in males.

Comments: Solier(1835) described T. subrugosa Dejan in litt. (Dejean 1821, 1833) from Spain, using a non-valid name by primary homonymy with T. subrugosa Besser, 1832, which was described with specimens from Calabria (Italy), collected by Parreiss (Parreys), and labelled as T. subrugosa Dejean. Thus, we propose T. circumvoluta nom. nov., replacing T. subrugosa Solier nom. preocc., by the circumvolutions of the elytral sculpture.

T. subrugosa Besser is an enigmatic species that has not been recorded since its description, partly due to the unawareness of the type material. Besser (1832) tried to accommodate it in the of Steven’s keys the specimens of “ T. subrugosa Dejean ” received from Parreys, including only those characters that he considers differentials with respect to other species like “ T. nomax, T. taurica and T. podolica ”, but that by themselves do not allow accurate identification. We have examined six specimens labelled “ Tentyria subrugosa Dej. Sicil. Parreys ” from NMHUB, but only two of them had old handwritten label (Fig. 259). These two specimens and another three belonging to the same taxa, are identical to T. italica Solier, 1835 (Fig. 260), which therefore becomes a synonym of T. subrugosa Besser, 1832; the sixth specimen corresponds to T. grossa Besser, 1832 (Fig. 261). Probably, the specimens deposited in the NMHUB are syntypes of T. subrugosa Besser and T. grossa Besser respectivamente (see previous comments referred to T. curculionoides Herbst and T. platyceps Steven).

Geographical distribution: Described from Spain without specifying locality. Apparently, it is present in the northern sub-plateau, in the foothills of the Central System. Very common in the province of Ávila (La Serrada, Español 1960); Viñolas & Cartagena (2005) also cited this species in Segovia; and Kraatz (1865) in Aranjuez and Madrid.