TribeAlfarini + Tribe Brachyrhaphini + Tribe Priapichthyini + Tribe Priapellini + Tribe Heterandriini + Tribe Gambusiini + Supertribe Poeciliini

[Clade 126]

Diagnosis. Members of this clade can be diagnosed by the following not uniquely derived and/or reversed features: (1) posterior supraorbital canal (2b, 3, 4a) opened, forming a sinuous depression over the frontal (supraorbital bone) [5-1*]; (2) anterior section of posterior remnant of infraorbital system (canal 4b, 5, 6a) opened, pores confluent forming a major sinuous depression above and slightly behind the orbit [6- 1]; (3) medial surface of ascending process of premaxilla slightly angled laterally [11-1*]; (4) anterior border of ventral maxilla concave [14-1]; (5) pelvic girdle of males posteriorly located; posterior border of cleithrum approximately aligned with center of basipterygium (or more posterior) [35-1]; (6) ligastyle with one axis [46-1]; and (7) adult males with anterior process on base of fifth middle anal-fin radial pointed and upward directed [78-1].